The concept of an ecological niche (econiche) has been used in a variety of ways, some of which are incompatible with a relational or functional interpretation of the term. This essay seeks to standardize usage by limiting the concept to functional relations between organisms and their surroundings, and to revise the concept to include epistemic relations. For most organisms, epistemics are a vital aspect of their functional relationships to their surroundings and, hence, a major determinant of their econiche. Rejecting the (...) traditional dualism of organism and environment, an econiche is defined as the reciprocal (dual) of a functionally specified class of organisms (FSTU). From this perspective, an econiche necessarily implies a certain type of organism, and a class of functionally similar organisms implies a special econiche.The econiche concept is also discussed in relation to other ecological terms that reflect the distributional patterns of organisms, such as habitat, and the concept of an empty niche is criticized. (shrink)

Are ecosystems morally considerable-that is, do we owe it to them to protect their “interests”? Many environmental ethicists, impressed by the way that individual nonsentient organisms such as plants tenaciously pursue their own biological goals, have concluded that we should extend moral considerability far enough to include such organisms. There is a pitfall in the ecosystem-to-organism analogy, however. We must distinguish a system’s genuine goals from the incidental effects, or byproducts, of the behavior of that system’s parts. Goals seem capable (...) of giving rise to interests; byproducts do not. It is hard to see how whole ecosystems can be genuinely goal-directed unless group selection occurs at the community level. Currently, mainstream ecological and evolutionary theory is individualistic. From such a theory it follows that the apparent goals of ecosystems are mere byproducts and, as such, cannot ground moral considerability. (shrink)

Wilderness, for seventeenth-century Puritan colonists in America, was hideous and howling. In the eighteenth century, Puritan preacher and theologian, Jonathan Edwards, began the process of transforming the American wilderness into an aesthetic and spiritual resource, a process completed in the nineteenth century by Ralph Waldo Emerson. Henry David. Thoreau was the first American to recommend wilderness preservation for purposes of transcendental recreation (solitude, and aesthetic and spiritual experience). In the twentieth century, Theodore Roosevelt and Aldo Leopold advocated wilderness preservation for (...) a different kind of recreation (hunting, fishing, and primitive travel) in order to preserve the putatively unique American character and institutions. Of these three historic conceptions of wilderness preservation, the third is the best model for frontier ecosystems at the austral tip of the Americas. (shrink)

Martha Nussbaum has expanded the capabilities approach to defend positive duties of justice to individuals who fall below Rawls’ standard for fully cooperating members of society, including sentient nonhuman animals. Building on this, David Schlosberg has defended the extension of capabilities justice not only to individual animals but also to entire species and ecosystems. This is an attractive vision: a happy marriage of social, environmental and ecological justice, which also respects the claims of individual animals. This paper asks whether it (...) is one that the capabilities approach can really deliver. Serious obstacles are highlighted. The potential for conflict between the capability-based entitlements of humans and those of nonhuman animals or ‘nature’ is noted, but it is argued that this does not constitute a decisive objection to the expanded capabilities approach. However, intra-nature conflicts are so widespread as to do so: the situation is outside the circumstances of justice as they are standardly understood. Schlosberg attempts to reconcile such conflicts by re-examining what it means to flourish as a sentient nonhuman animal. This fails, because of the distinction between flourishing as a species, which often requires predation, and flourishing as an individual, which is as frequently incompatible with it. Finally, the paper considers how a capabilities theorist might move beyond such conflicts, identifying two possible strategies (which are not themselves unproblematic) for reconciling the demands of humans, animals and ecosystems. (shrink)

In 1935, A. G. Tansley, who was knighted later, proposed the ecosystem concept. Nevertheless, this concept was not without predecessors. Why did Tansley’s ecosystem prevail and not one of its competitors? The purpose of this article is to pin the distinguishing features of Tansley’s ecosystem down, as far as the published record allows. It is an exercise in finding the difference that made a difference. Besides being a pioneering ecologist, Tansley was an adept of psychoanalysis. His interest even led him (...) to visit Sigmund Freud in Vienna for a while. Psychologists had to regard the mind as an entity in its own right, while knowing that it truly was part of a larger whole (body + mind), because the causal relation between body and mind was unknown. This lead Tansley to conclude that psychologists must not objectify the system under study, have to search for causes within their own field, and must not speculate unless this serves a scientific purpose. In 1925, Tansley defended psychoanalysis in a prolonged controversy against a concerted attack criticizing its speculative content and poor scientific standing. This could have been the reason why Tansley kept his ecosystem free of speculative content and unscientific connotation. The competing ecosystem-like concepts, however, have contained philosophical speculation, non-deterministic properties like vitalism or entelechy, or have been burdened with unscientific connotations. Hence, rigorous restraint distinguished the ecosystem concept and made it ready for use by later researchers. (shrink)

Philosophers of the life sciences have devoted considerably more attention to evolutionary theory and genetics than to the various sub-disciplines of ecology, but recent work in the philosophy of ecology suggests reflects a growing interest in this area (Cooper 2003; Ginzburg and Colyvan 2004). However, philosophers of biology and ecology have focused almost entirely on conceptual and methodological issues in population and community ecology; conspicuously absent are foundational investigations in ecosystem ecology. This situation is regrettable. Ecosystem concepts play a central (...) role in many branches of theoretical and applied ecology, and in environmental literature generally. Indeed, for some historians, the division of ecological theory into population-community and ecosystem research traditions, and the methodological and conceptual debates that have arisen between workers in these respective camps, is the distinguishing feature of 20th century ecological science (Hagen 1992). These include debates over, among others: reductionistic vs. holistic research methodologies; the existence and metaphysics of ecological “kinds”; the relationship between evolutionary mechanisms and ecosystem phenomena; and the nature and scope of ecological science and its relationship to other branches of natural and social science. Philosophers of ecology have written on all these topics, but almost exclusively from the theoretical perspective of population, community or evolutionary ecology. Philosophical attention to these issues from the perspective of ecosystem ecology is long overdue. It would be misleading to assert that philosophers in general have ignored ecosystem ecology. Environmental philosophers, including environmental ethicists so-called “radical” environmental philosophers (deep ecologists, social ecologists, ecofeminists, etc.), and policy theorists, have had a long-standing interest in ecosystem ecology (e.g. Callicott 1986; Cahen 1988; Warren and Cheney 1993; Westra 1994; Sagoff 1997; Fitzsimmons 1999).. (shrink)

Environmental ethics is generally searching for the intrinsic value in natural beings. However, there are very few holistic models trying to reflect the various dimensions of the experience-to-be a natural being. We are searching for that intrinsic value, in order to determine which species are holders of rights. In this article, I suggest a set of moral and rational principles to be used for identifying the intrinsic value of a given species and for comparing it to that of other species.

The world is an untidy place, and the sciences—all of them—reflect this. One source of this untidiness is the relationship between levels of organization. Reducing macrolevels to microlevels—explaining the former in terms of the latter—has met with successes but has never been the whole story. In the biological sciences, there has been much attention lately to the shortcomings of reductionism on the grounds that (i) it changes the subject rather than explaining, (ii) it leads to a myopically molecular view of (...) the biological world, and (iii) the behavior or behaviors of complex systems are often very poorly predicted based solely on their microproperties. It is just for these reasons that biologists of many stripes have called for a move away from reductionism and toward a new kind of biology for the 21st century. But what shape might this new biology take? (shrink)

Sound definitions of its basic concepts are fundamental to every scientific discipline. In some instances, like in the case of the ecosystem concept, the question arises if we can define such concepts at all. And if we can define them, how should we choose from the multiple definitions available? And what are the preconditions for a scientifically sound and useful definition? On the basis of the ecosystem concept, this paper illustrates a major, often neglected distinction in the definition of ecological (...) concepts, namely that between defining criteria and additional descriptive statements connected to those definitions. As is demonstrated by examples from the literature, mixing up these categories leads to false inferences about the properties of physical objects (e.g. a particular forest) subsumed by the concept (e.g. the ecosystem). As a further consequence, this inference becomes problematic in terms of theory development and/or the application of ecological concepts for management decisions. (shrink)

I develop the thesis that species and ecosystems are living entities with morally significant interests in their own right and defend it against leading objections. Contrary to certain claims, it is possible to individuate such entities sufficiently well. Indeed, there is a sense in which such entities define their own nature. I also consider and reject the argument that species and ecosystems cannot have interests or even traits in their own right because evolution does not proceed on that level. Although (...) evolution proceeds on the level of the genotype, those selected are able to cooperate in entities of various higher orders—including species and ecosystems. Having their own nature and interests, species and ecosystems can meaningfully be said to have moral standing. (shrink)

A formal theory of the ecological niche is indispensable not only for semantic precision in philosophy to understand and compare it with other meanings of niche, but also when computer scientists and ecologists desire to create interoperable software where one can retrieve the niche of a species and compare their parameters. The proposed model is a more fine-grained description of the ecological niche, including the distinction between its complex concept, the abstract niche (‘fundamental niche’) with its hypervolume in multidimensional space, (...) and its realisations (‘realised niches’). The presented ecological niche may initiate new avenues for research in ecology, particularly concerning the conditions/categories of a hypervolume, as well as further philosophical inquiry and comparison with other niches. (shrink)

The dynamics of single populations up to ecosystems, are often described by one or a set of non-linear ordinary differential equations. In this paper we review the use of bifurcation theory to analyse these non-linear dynamical systems. Bifurcation analysis gives regimes in the parameter space with quantitatively different asymptotic dynamic behaviour of the system. In small-scale systems the underlying models for the populations and their interaction are simple Lotka-Volterra models or more elaborated models with more biological detail. The latter ones (...) are more difficult to analyse, especially when the number of populations is large. Therefore for large-scale systems the Lotka-Volterra equations are still popular despite the limited realism. Various approaches are discussed in which the different time-scale of ecological and evolutionary biological processes are considered together. (shrink)

Abstract Traditional moral philosophy, developmental psychology and moral education have generally been concerned with relationships between human beings. However, moral philosophy has gradually expanded to include plants, animals and ecosystems as legitimate moral objects, and aesthetics has rediscovered nature as an object of consideration. Thus it seems appropriate to begin to include this sphere in moral education and corresponding research as well. In this paper we wish to report on an investigation we have begun using children's philosophy as a hermeneutic (...) tool to assess the morally relevant values and attitudes that children and adolescents hold with respect to nature and the patterns of reasoning with which they are expressed. In the first part of our presentation we will outline current positions in environmental ethics and aesthetics which provide a theoretical framework for such an investigation and the problems they present. Subsequently the difficulties involved with applying contemporary theories of moral development to the problem at hand will be discussed. Finally, we will describe the project we have begun and summarise preliminary results derived from different age groups. (shrink)

Philosophers of the life sciences have devoted considerably more attention to evolutionary theory and genetics than to the various sub-disciplines of ecology, but recent work in the philosophy of ecology suggests reflects a growing interest in this area (Cooper 2003; Ginzburg and Colyvan 2004). However, philosophers of biology and ecology have focused almost entirely on conceptual and methodological issues in population and community ecology; conspicuously absent are foundational investigations in ecosystem ecology. This situation is regrettable. Ecosystem concepts play a central (...) role in many branches of theoretical and applied ecology, and in environmental literature generally. Indeed, for some historians, the division of ecological theory into population-community and ecosystem research traditions, and the methodological and conceptual debates that have arisen between workers in these respective camps, is the distinguishing feature of 20th century ecological science (Hagen 1992). These include debates over, among others: reductionistic vs. holistic research methodologies; the existence and metaphysics of ecological “kinds”; the relationship between evolutionary mechanisms and ecosystem phenomena; and the nature and scope of ecological science and its relationship to other branches of natural and social science. Philosophers of ecology have written on all these topics, but almost exclusively from the theoretical perspective of population, community or evolutionary ecology. Philosophical attention to these issues from the perspective of ecosystem ecology is long overdue. It would be misleading to assert that philosophers in general have ignored ecosystem ecology. Environmental philosophers, including environmental ethicists so-called “radical” environmental philosophers (deep ecologists, social ecologists, ecofeminists, etc.), and policy theorists, have had a long-standing interest in ecosystem ecology (e.g. Callicott 1986; Cahen 1988; Warren and Cheney 1993; Westra 1994; Sagoff 1997; Fitzsimmons 1999).. (shrink)

In this essay I first provide an analysis of various community concepts. Second, I evaluate two of the most serious challenges to the existence of communities—gradient and paleoecological analysis respectively—arguing that, properly understood, neither threatens the existence of communities construed interactively. Finally, I apply the same interactive approach to ecosystem ecology, arguing that ecosystems may exist robustly as well. ‡I would like to thank to the participants at the Ecology and Environmental Ethics Conference at the University of Utah, the Philosophy (...) of Ecology Conference hosted by the University of Brisbane, and those participants in a session at the Philosophy of Science Association Meeting in Vancouver, British Columbia for helpful discussions of this essay. Specific thanks go to Mark Colyvan, Greg Cooper, Steve Downes, Chris Elliott, Marc Ereshefsky, Paul Griffiths, Jesse Hendrikse, Greg Mikkelson, Anya Plutynski, Kate Ritchie, Sahotra Sarkar, Kim Sterelny, and Rob Wilson. †To contact the author, please write to: Department of Philosophy, Lewis and Clark College, 0615 SW Palatine Hill Road, Portland, OR 97219; e-mail: jay@lclark.edu. (shrink)