Four notions of biological function

Abstract
I argue that there are at least four different ways in which the term 'function' is used in connection with the study of living organisms, namely: (1) function as (mere) activity, (2) function as biological role, (3) function as biological advantage, and (4) function as selected effect. Notion (1) refers to what an item does by itself; (2) refers to the contribution of an item or activity to a complex activity or capacity of an organism; (3) refers to the value for the organism of an item having a certain character rather than another; (4) refers to the way in which a trait acquired and has maintained its current share in the population. The recognition of a separate notion of function as biological advantage solves the problem of the indeterminate reference situation that has been raised against a counterfactual analysis of function, and emphasizes the importance of counterfactual comparison in the explanatory practice of organismal biology. This reveals a neglected problem in the philosophy of biology, namely that of accounting for the insights provided by counterfactual comparison.
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Helena Siipi (2008). Dimensions of Naturalness. Ethics and the Environment 13 (1):pp. 71-103.
Thomas A. C. Reydon (2009). Gene Names as Proper Names of Individuals: An Assessment. British Journal for the Philosophy of Science 60 (2):409-432.

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