Online research in philosophy

The concept of niche (setting, context, habitat, environment) has been little studied by ontologists, in spite of its wide application in a variety of disciplines from evolutionary biology to economics. What follows is a first formal theory of this concept, a theory of the relations between objects and their niches. The theory builds upon existing work on mereology, topology, and the theory of spatial location as tools of formal ontology. It will be illustrated above all by means of simple biological examples, but the concept of niche should be understood as being, like concepts such as part, boundary, and location, a structural concept that is applicable in principle to a wide range of different domains.

The concept of niche (setting, context, habitat, environment) has been little studied by ontologists, in spite of its wide application in a variety of disciplines from evolutionary biology to economics. What follows is a first formal theory of this concept, a theory of the relations between objects and their niches. The theory builds upon existing work on mereology, topology, and the theory of spatial location as tools of formal ontology. It is illustrated above all by means of simple biological examples, but the concept of niche should be understood as being, like concepts such as part, boundary, and location, a structural concept that is applicable in principle to a wide range of different domains.

Industrial melanism, according to the traditional explanation, amounts to niche construction since it involves changes in predation pressure. Indeed, it would be difficult to imagine selection without niche construction. This cannot be what Laland, Odling-Smee & Feldman mean. They offer convincing examples, but they should provide a better definition of “niche construction” to indicate how their view supplements traditional evolutionary biology.

The concept of an ecological niche (econiche) has been used in a variety of ways, some of which are incompatible with a relational or functional interpretation of the term. This essay seeks to standardize usage by limiting the concept to functional relations between organisms and their surroundings, and to revise the concept to include epistemic relations. For most organisms, epistemics are a vital aspect of their functional relationships to their surroundings and, hence, a major determinant of their econiche. Rejecting the traditional dualism of organism and environment, an econiche is defined as the reciprocal (dual) of a functionally specified class of organisms (FSTU). From this perspective, an econiche necessarily implies a certain type of organism, and a class of functionally similar organisms implies a special econiche.The econiche concept is also discussed in relation to other ecological terms that reflect the distributional patterns of organisms, such as habitat, and the concept of an empty niche is criticized.

Recent theories in cognitive science have begun to focus on the active role of organisms in shaping their own environment, and the role of these environmental resources for cognition. Approaches such as situated, embedded, ecological, distributed and particularly extended cognition look beyond ‘what is inside your head’ to the old Gibsonian question of ‘what your head is inside of’ and with which it forms a wider whole—its internal and external cognitive niche. Since these views have been treated as a radical departure from the received view of cognition, their proponents have looked for support to similar extended views within (the philosophy of) biology, most notably the theory of niche construction. This paper argues that there is an even closer and more fruitful parallel with developmental systems theory and developmental niche construction. These ask not ‘what is inside the genes you inherited’, but ‘what the inherited genes are inside of’ and with which they form a wider whole—their internal and external ontogenetic niche, understood as the set of epigenetic, social, ecological, epistemic and symbolic legacies inherited by the organism as necessary developmental resources. To the cognizing agent, the epistemic niche presents itself not just as a partially self-engineered selective niche, as the niche construction paradigm will have it, but even more so as a partially self-engineered ontogenetic niche, a problem-solving resource and scaffold for individual development and learning. This move should be beneficial for coming to grips with our own (including cognitive) nature: what is most distinctive about humans is their developmentally plastic brains immersed into a well-engineered, cumulatively constructed cognitive–developmental niche.

This article examines some of the main tenets of competition theory in light of the theory of evolution and the concept of an ecological niche. The principle of competitive exclusion and the related assumption that communities exist at competitive equilibrium - fundamental parts of many competition theories and models - may be violated if non-equilibrium conditions exist in natural communities or are incorporated into competition models. Furthermore, these two basic tenets of competition theory are not compatible with the theory of evolution. Variation in ecologically significant environmental factors and non-equilibrium in population numbers should occur in most natural communities, and such changes have important effects on community relations, niche overlap, and the evolution of ecosystems. Ecologists should view competition as a process occurring within a complexdynamic system, and should be wary of theoretical positions built upon simple laboratory experiments or simplistic mathematical models.In considering the relationship between niche overlap and competition, niche overlap should not be taken as a sufficient condition for competition; many factors may prevent or diminish competition between populations with similar resource utilization patterns. The typically opposing forces of intraspecific and interspecific competition need to be simultaneously considered, for it is the balance between them that in large part determines niche boundaries.

Some species are weedy: they move from one ecological niche to another. Other species are specialized: they are exquisitely adapted to exploit a particular niche. Human beings are the design solution in which a species is simultaneously weedy and specialized - the trick being to manage the exquisite niche-specific adaptations in software rather than in the hardware. We are built to reprogram ourselves on the fly, to select new goals, new priorities and new guidelines appropriate to novel niches. Understanding ourselves as an implementation of this design solution has consequences for the theory of practical reasoning. Instrumentalism (the theory of practical reasoning according to which it consists solely in selecting means to pre-given ends) cannot be a suitable theory of rationality for such a species (that is, our own).

For several decades, the concept of the niche has enriched sociological theory. The niche represents the position or function of an entity, such as an organization or population of organizations, within a larger community environment. Using the concept of the niche allows researchers to go beyond classifying entities to understanding (a) their life chances under different and changing environmental conditions and (b) how they interact under the competitive conditions induced by a finite environment. We briefly review the intellectual history of the niche concept as it came from bioecology to sociology. The bulk of the article reviews the two major streams of sociological research that use the niche concept: the population ecology of organizations and McPherson's ecology of affiliation. Finally, we survey commonalities between these approaches and highlight new directions, including applications that take the niche concept well beyond its origins in organizational studies.

Although the concept of niche has been extremely useful in sociological theory and research, some aspects of the concept have not been clearly developed. This article advances a theoretical reconstruction of the concept of niche, with special application to organizations. The proposed formal model unifies several active lines of sociological theory. It also extends the notion of the niche from the realm of behaviors to apply to the rules coding social identities and organizational forms. The reconstruction gives deeper insight into the niche of an organizational population as well as individual organizations. Finally, the model analyzes the (thus far) tacit assumption that niches are convex, examines the implications of convexity for commonly used measures of niche width, and provides a general sociological argument for the predominance of convex niches.

The ecological literature distinguishes between two ways of conceiving a “niche” (habitat, ecotope, biotope, microlandscape) [22, 39]. On the one hand, there is the traditional functional conception of a niche as the role or position enjoyed by an organism or population within an ecological community. As C. Elton [14] famously put it, “When an ecologist says ‘there goes a badger’ he should include in his thoughts some definite idea of the animal’s place in the community to which it belongs, just as if he had said ‘there goes the vicar’.” The world of niches might, in this sense, be viewed as a giant evolutionary hotel, some of whose rooms are occupied (by organisms which have evolved to fill them), some of whose rooms are for a variety of reasons unoccupied but can become occupied in the future. On the other hand, there is the environmental conception advanced by G. E. Hutchinson [19] and R. Lewontin [21]. On this second conception, a niche is thought of as the hypervolume defined by the limiting values of all environmental variables relevant to the survival of a given species. A niche is not a mere location, but a location in space that is defined additionally by a specific constellation of ecological parameters such as degree of slope, exposure to sunlight, soil fertility, foliage density, and so on. It is, we might say, an ecological context. The purpose of this paper is to outline a formal theory of this notion. Our account expands on the theory put forward in [30], which builds upon certain fundamental notions and principles of mereology, topology, and the theory of spatial location. We focus on niche tokens, which is to say on the environmental niche determined by a given organism or population of organisms in a given place, and we aim to be more explicit than is customary in the ecological literature as concerns the..