Screening-off and the units of selection

Philosophy of Science 59 (1):142-152 (1992)
Brandon ([1982] 1984, 1990) has argued that Salmon's (1971) concept of screening-off can be used to characterize (i) the idea that natural selection acts directly on an organism's phenotype, only indirectly on its genotype, and (ii) the biological problem of the levels of selection. Brandon also suggests (iii) that screening-off events in a causal chain are better explanations than the events they screen off. This paper critically evaluates Brandon's proposals
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    James F. Crow (1994). In Praise of Replicators. Behavioral and Brain Sciences 17 (4):616.

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