The diversity, complexity and adaptation of the biological realm is evident. Until Darwin, the best explanation for these three features of the biological was the conclusion of the “argument from design.” Darwin's theory of natural selection provides an explanation of all three of these features of the biological realm without adverting to some mysterious designing entity. But this explanation's success turns on the meaning of its central explanatory concept, ‘fitness’. Moreover, since Darwinian theory provides the resources for a purely causal (...) account of teleology, wherever it is manifested, its reliance on the concept of ‘fitness’ makes it imperative that conceptual problems threatening the explanatory legitimacy of this notion be solved. (shrink)

Recent discussions in the philosophy of biology have brought into question some fundamental assumptions regarding evolutionary processes, natural selection in particular. Some authors argue that natural selection is nothing but a population-level, statistical consequence of lower-level events (Matthen and Ariew [2002]; Walsh et al. [2002]). On this view, natural selection itself does not involve forces. Other authors reject this purely statistical, population-level account for an individual-level, causal account of natural selection (Bouchard and Rosenberg [2004]). I argue that each of these (...) positions is right in one way, but wrong in another; natural selection indeed takes place at the level of populations, but it is a causal process nonetheless. (shrink)

Applications of game theory to moral philosophy are impededby foundational issues and troublesome examples. In the first part of this paper,questions are raised about the appropriate game-theoretical frameworks for applications to moralphilosophy and about the proper interpretations of the theoretical devices employed inthese frameworks. In the second part, five examples that should be of particular interest to thoseinterested in the connections between ethics and game theory are delineated and discussed. Thefirst example comprises games in which there is an outcome unanimously (...) preferred to the``solution'' of the game, appropriately defined. The second comprises games whose solution callsfor different players to employ different strategies. The third comprises games whosesolution calls for players to adopt mixed strategies. The fourth comprises games whose solutionrequires players to cycle among a variety of strategies. The fifth comprises games whose solutionrequires players to discriminate in morally inappropriate ways. (shrink)

The development of evolutionary game theory is closely linked with two interdisciplinary exchanges: the import of game theory into biology, and the import of biologists’ version of game theory into economics. This paper traces the history of these two import episodes. In each case the investigation covers what exactly was imported, what the motives for the import were, how the imported elements were put to use, and how they related to existing practices in the respective disciplines. Two conclusions emerged from (...) this study. First, concepts derived from the unity of science discussion or the unification accounts of explanation are too strong and too narrow to be useful for analysing these interdisciplinary exchanges. Secondly, biology and economics—at least in relation to EGT—show significant differences in modelling practices: biologists seek to link EGT models to concrete empirical situations, whereas economists pursue conceptual exploration and possible explanation.Keywords: Models; Evolutionary game theory; Interdisciplinarity; Unification; Unity of science; Theory import; Biology; Economics. (shrink)

Both biologists and philosophers often make use of simple verbal formulations of necessary and sufficient conditions for evolution by natural selection (ENS). Such summaries go back to Darwin's Origin of Species (especially the "Recapitulation"), but recent ones are more compact.1 Perhaps the most commonly cited formulation is due to Lewontin.2 These summaries tend to have three or four conditions, where the core requirement is a combination of variation, heredity, and fitness differences. The summaries are employed in several ways. First, they (...) are often used in pedagogical contexts, and in showing the coherence of evolutionary theory in response to attacks from outside biology. Second, they are important in discussions of extensions of evolutionary principles to new domains, such as cultural change. The summaries also have intrinsic scientific and philosophical interest as attempts to capture some core principles of evolutionary theory in a highly concise way. Despite their prominence, both the proper formulation and status of these summaries are unclear. Standard formulations are subject to counterexamples, and their relations to formal models of evolutionary change are not straightforward. Here I look closely at these verbal summaries, and at how they relate to formal models. Are the summaries merely rough approximations that have no theoretical role of their own? Perhaps they could operate as theoretical statements in Darwin's time, but have now been superseded by more exact treatments. (shrink)