The semantic claim defended in this article is that the Participle II morphology is not linked to a uniform meaning. The meaning rather co-varies with the syntactic function of the participle. As..
Philosophers differ widely in the extent to which they condone the exploration of the realms of possibilia. Some are very enamoured of thought experiments in which human intuition is trained upon the products of human imagination. Others are much more sceptical of the fruits of such purely cognitive explorations. That said, it is clear that human beings cannot dispense with modal speculation altogether. Rationality rests upon the ability to make decisions and that in turn rests upon the ability to learn (...) about what is possible and what is probable. Thus, on pain of irrationality, we must have some means of exploring other possible worlds. Thankfully, intuition is not the only aid we have at our disposal. Science also is in the business of finding regularities, which hold counterfactually. Scientific theory tells us about the likelihood of particular outcomes flowing from particular processes given particular background conditions. Thus, it also tells us about the contents of 1 of 21 other possible worlds. One consequence of the possibility of such inferences has been a theoretical interest, not just in the contents, but also in the geography of the domain of all possibly worlds. Metaphysicians, epistemologists and philosophers of language are very familiar with locutions such as “nearby possible worlds” (meaning possible worlds very similar to the actual world). Similarly, evolutionary theory tells us that there is little chance of us discovering an organism that is mammal-like in most respects except in having six limbs. It’s not that we know such an organism to be impossible, but rather that we think it would be the product of an evolutionary history very different to the actual history of life on earth. Put another way, such organisms would be denizens of distant possible worlds. Clearly then, both biology and philosophy have ample motivation to be interested in the reasoning and evidence that supports such claims. Seemingly, in both disciplines there is a certain lure to this modal cartography, but ought we in fact to be convinced of its merits? Is it science or philosophy or not a good example of either? What sort of problems can it solve? What sort of problems will it create? How might we test its accuracy? In his excellent book Theoretical Morphology: The Concept and Its Applications (1999), George McGhee provides an admirable introduction to the complex theoretical landscape surrounding the exploration of possible biological form.. (shrink)
In plant morphology, most structures of vascular plants can easily be assigned to pre-established organ categories. However, there are also intermediate structures that do not fit those categories associated with a classical approach to morphology. To integrate the diversity of forms in the same general framework, we constructed a theoretical morphospace based on a variety of modalities where it is possible to calculate the morphological distance between plant organs. This paper gives emphasis on shoot, leaf, leaflet and trichomes (...) while ignoring the root. This will allow us to test the hypothesis that classical morphology (typology) and dynamic morphology occupy the same theoretical morphospace and the relationship between the two approaches remains a question of weighting of criteria. Our approach considers the shoot (i.e. leafy stem) as the basic morphological structural unit. A theoretical data table consisting of as many lines as there are possible combinations between different modalities of characters of a typical shoot was generated. By applying a principal components analysis (PCA) to these data it is possible to define a theoretical morphospace of shoots. Typical morphological elements (shoots, leaves, trichomes) and atypical structures (phylloclades, cladodes) including particular cases representing ‘exotic’ structures such as the epiphyllous appendages of Begonia and ‘water shoot’ and ‘leaf’ of aquatic Utricularia were placed in the morphospace. The more an organ differs from a typical shoot, the further away it will be from the barycentre of shoots. By giving a higher weight to variables used in classical typology, the different organ categories appear to be separate, as expected. If we do not make any particular arbitrary choice in terms of character weighting, as it is the case in the context of dynamic morphology, the clear separation between organs is replaced by a continuum. Contrary to typical structures, “intermediate” structures are only compatible with a dynamic morphology approach whether they are placed in the morphospace based on a ponderation compatible with typology or dynamic morphology. The difference in points of view between typology and continuum leads to a particular mode of weighting. By using an equal weighting of characters, contradictions due to the ponderation of characters are avoided, and the morphological concepts of continuum’ and ‘typology’ appear as sub-classes of ‘process’ or ‘dynamic morphology’. (shrink)
One foundational question in contemporarybiology is how to `rejoin evolution anddevelopment. The emerging research program(evolutionary developmental biology or`evo-devo) requires a meshing of disciplines,concepts, and explanations that have beendeveloped largely in independence over the pastcentury. In the attempt to comprehend thepresent separation between evolution anddevelopment much attention has been paid to thesplit between genetics and embryology in theearly part of the 20th century with itscodification in the exclusion of embryologyfrom the Modern Synthesis. This encourages acharacterization of evolutionary developmentalbiology as the marriage (...) of evolutionary theoryand embryology via developmental genetics. Butthere remains a largely untold story about thesignificance of morphology and comparativeanatomy (also minimized in the ModernSynthesis). Functional and evolutionarymorphology are critical for understanding thedevelopment of a concept central toevolutionary developmental biology,evolutionary innovation. Highlighting thediscipline of morphology and the concepts ofinnovation and novelty provides an alternativeway of conceptualizing the `evo and the `devoto be synthesized. (shrink)
This paper offers an account of the relationship between inference and explanation in functional morphology which combines Robert Brandon's theory of adaptation explanation with standard accounts of inference to the best explanation. Inferences of function from structure, it is argued, are inferences to the best adaptation explanation. There are, however, three different approaches to the problem of determining which adaptation explanation is the best. The theory of inference to the best adaptation explanation is then applied to a case study (...) from the history of functional morphology: the case of the crested duckbilled dinosaurs. (shrink)
With Carl Gegenbaur and Ernst Haeckel, inspiredby Darwin and the cell theory, comparativeanatomy and embryology became established andflourished in Jena. This tradition wascontinued and developed further with new ideasand methods devised by some of Haeckelsstudents. This first period of innovative workin evolutionary morphology was followed byperiods of crisis and even a disintegration ofthe discipline in the early twentieth century.This stagnation was caused by a lack ofinterest among morphologists in Mendeliangenetics, and uncertainty about the mechanismsof evolution. Idealistic morphology was (...) stillinfluental in Germany, which prevented a fullappreciation of the importance of Darwinstheory of natural selection for comparativemorphology. Evolutionary morphology andembryology failed to contribute significantlyto the modern synthesis of evolutionarybiology, thereby probably delaying theintegration of developmental biology intomodern evolutionary biology. However, Haeckelsstudent Oscar Hertwig, as well as Victor Franzand Alexej N. Sewertzoff from a youngergeneration, all tried to forge their ownsynthetic approaches in which (inspired byHaeckels work) embryology played an importantrole. Important for all three researchers wereattempts to refine, and sometimes redefine, thebiogenetic law, and to find new scientificexplanations for it (and for the manyexceptions to it). Their research was latermore or less forgotten, and had littleinfluence on the architects of the modernsynthesis. As the relationship betweenevolutionary and developmental biology is nowagain rising in importance in the form ofEvo-Devo, we would like to draw attention tohow this earlier research tradition grappledwith similar questions to those now on theagenda, albeit from sometimes quite differentperspectives. (shrink)
One foundational question in contemporary biology is how to integrate evolution and development. The emerging synthesis (evolutionary developmental biology or ‘evo-devo’) requires a meshing of disciplines, concepts, and explanations (inter alia) that have been developed largely in independence over the past century. The nature of the hoped for synthesis is not wholly agreed upon due to divergent viewpoints resulting from this disciplinary independence and, consequently, the mechanics for accomplishing the task are not clearly specified. This paper utilizes historical investigation for (...) philosophical purposes in order to explore the question of synthesizing evolutionary and developmental biology. In the attempt to comprehend the present separation between evolution and development much attention has been paid to the split between genetics and embryology in the early part of the century with its codification in the exclusion of embryology from the Modern Synthesis. This encourages a characterization of "evo-devo" as the integration of developmental genetics with Neo-Darwinism. But there is a largely untold story about the significance of morphology and comparative anatomy (also minimized in the Modern Synthesis). I will attempt to reconstruct part of this story, focusing on the rebirth of functional (and evolutionary) morphology after the 1950s. Functional morphology is critical for understanding the development of a concept central to "evo-devo", evolutionary innovation. Understanding the story about morphology and innovation reveals a different conception of the foundational problem, providing alternative ways of conceptualizing the "evo" and the "devo" to be synthesized. (shrink)
This paper examines a new challenge to neo-Darwinism, a movement known as process structuralism. The process structuralist critique of neo-Darwinism holds 1) that there are general laws in biology and that biologists should search for these laws; 2) that there are general forms of morphology and development and that biologists should attempt to uncover these forms; 3) that organisms are unified wholes that cannot be understood without adopting a holistic perspective; and 4) that no special, causal primacy should be (...) given to the genes in development and morphology. This paper places process structuralism in its historical context, examines the philosophical underpinnings of the movement, and discusses some of the evidence used to support its claims. (shrink)
From the point of view of a dynamic morphology, form is not only the result of process(es) — it is process. This process may be analyzed in terms of two pairs of fundamental processes: growth and decay, differentiation and dedifferentiation. Each of these processes can be analyzed in terms of various modalities (parameters) and submodalities. This paper deals with those of growth (see Table 1). For the purpose of systematits and phylogenetic reconstruction the modalities and submodalities can be considered (...) dynamic characters that have states. Each state of such a dynamic character is a more detailed process, hence not static. For example, determinate growth represents a state of the dynamic character (or modality) of growth duration.The processes of Table 1 can be applied to the whole plant kingdom (although in certain cases only some processes of the whole set may be applicable). Thus, the diversity of plant form is seen as a diversity of process combinations. From this point of view, change in form implies change in the process combination(s). Questions that arise are, for example, the following: Which process combinations actually occur? Which of these are the most frequent? How and why have process combinations changed during ontogeny and phylogeny? (shrink)
Evolutionary change is opportunistic, but its course is strongly constrained in several fundamental ways. These constraints (historical/phylogenetic, functional/adaptive, constructional/morphogenetic) and their dynamic relationships are discussed here and shown to constitute the conceptual framework of Constructional Morphology. Notwithstanding recent published opinions which claim that the discovery of constraints renders Neodarwinian selection theory obsolete, we regard the insights of Constructional Morphology as being entirely consistent with this theory. As is shown here in the case of the Hyracoidea, formal analysis of (...) the constraints which have framed the evolution of various characters extends our understanding of the evolution of a taxon. (shrink)
This paper is about a general methodology for pattern transformation. Patterns are network representations of the relations among structures and functions within an organism. Transformation refers to any realistic or abstract transformation relevant to biology, e.g. ontogeny, evolution and phenotypic clines. The main aim of the paper is a methodology for analyzing the range of effects on a pattern due to perturbing one or more of its structures and/or functions (transformation morphology). Concepts relevant to such an analysis of pattern (...) transformation are reviewed and several new ones introduced: pattern unit; direct and indirect functional demands; compatibility and trade-off; integrating, adding and decoupling; functional effectiveness; spatial, profile and other architectonic constraints; domains of structure-function relations; goal and process adaptability; multiple pathways. The paper is written from the the perspective of architectonic morphology, viz. functional morphology focusing on the relation between anatomical coherence and the compatibility of functions. The advantages and disadvantages of inductive and deductive approaches are discussed. (shrink)
Richard Owen has been condemned by Darwinians as an anti-evolutionist and an essentialist. In recent years he has been the object of a revisionist analysis intended to uncover evolutionary elements in his scientific enterprise. In this paper I will examine Owen's evolutionary hypothesis and its connections with von Baer's idea of divergent development. To give appropriate importance to Owen's evolutionism is the first condition to develop an up-to-date understanding of his scientific enterprise, that is to disentagle Owen's contribution to the (...) modernization of typology and morphology. I will argue that Owen's Platonic essentialism is rhetorical and incongruous. On the contrary, an interpretation of the archetype based on Aristotle's biological works makes possible a new conception of type, based on a homeostatic mechanism of stability. The renewal of morphology hinges on homological correspondences and a homeostatic process is also the origin of serial and special homology. I will argue that special homology shows an evolutionary orientation insofar as it is a typically inter-specific character while serial homology is determined through an elementary usage of the categories of developmental morphology. (shrink)
Analogy plays an important role in the production of irregular forms but the proposed Minimalist Morphology (MM) representations do not express this. Recent results also show that the regular forms of strong paradigms can have idiosyncratic properties that cannot be accounted for by MM. Methodological problems with an experiment are discussed and a plea for a processing explanation is made.
Plants are interpreted as structural hierarchies which are real systems organized through descending constraints. Types of hierarchical groups in plants are (a) cluster by integration, (b) support through attachment, (c) enclosure by encasement (d) dissipative by input of energy and (e) control through variable state switching. Most plant hierarchies are mixtures of these types which explains a number of paradoxes in plant morphology. The traditional means of identifying levels, i.e., cell, tissues, organs, uses a compositional group which is not (...) a hierarchical group but a similarity feature and so is inadequate for describing hierarchies. Hierarchies can be defined by set theory which is more a description of cognitive than real hierarchies and therefore is of little value in describing plant organization. The hierarchical description of a plant emphasizes the immediate physical status of organization which provide, in turn, a physical explanation of development. (shrink)
Notwithstanding the general rise of experimental disciplines in biology in the first decades of our century, in Germany and in the Netherlands the interest in the idealistic morphological tradition flourished, and compensated for a reductionistic causal approach to natural phenomena. This article analyses the influence of the German idealistic morphologists W. Lubosch and A. Meyer on the development of C.J. van der Klaauw's epistemology. It discusses the gradual incorporation of non-causal principles into van der Klaauw's concept of biology. Van der (...) Klaauw's epistemological concept of holistic biology was shaped in a critical confrontation with German idealistic morphology, and his early considerations can be interpreted as a direct impulse towards the development of his theory of functional components. Van der Klaauw's theories, being an alternative to the reductionistic experimental sciences, were among the causes of the fact that in the first half of our century biology in the Netherlands took a course deviating from the development of biology in the Anglo-American countries. (shrink)
The precise relationship between callosal morphology and specific connectivity is not yet known. Callosal axons are often presumed to be arranged according to their origin. In humans, this is true for the genu and the splenium, which convey axons from the prefrontal and occipital cortices, respectively, but not for the body, where axons from wide parts of the cortex are intermingled.
This paper shows a full description of Spanish inflectional morphology. We have chosen a paradigmatic approach instead of one based on phonological/spelling changes, i.e., the typical two-level model. Such morphological description has been written in the DATR formalism. The result is a network of nodes that makes use of the information inheritance mechanisms – orthogonal node inheritance and default path inheritance – that DATR allows. Some lexical coverage and corpus occurrence figures that support our approach are also given.
Handaxe morphology is thought to be the first example of the imposition of arbitrary form. Handaxes may thus inform researchers about shared mental templates and evolving cognitive abilities. However, many factors, not related to changes in cognition (e.g., material type, function, resharpening processes), influence handaxe shape over time and space. Archaeologists must control for these factors before making inferences concerning cognition.
Classical temples in ancient Greece show two deterministic illusionistic principles of architecture, which govern their functional design: geometric proportionalism and a set of illusion-strengthening rules in the proportionalism's stochastic margin. Animal morphology, in its mechanistic-deductive revival, applies just one architectural principle, which is not always satisfactory. Whether a Greek Classical situation occurs in the architecture of living structure is to be investigated by extreme testing with deductive methods.Three deductive methods for explanation of living structure in animal morphology are (...) proposed: the parts, the compromise, and the transformation deduction. The methods are based upon the systems concept for an organism, the flow chart for a functionalistic picture, and the network chart for a structuralistic picture, whereas the optimal design serves as the architectural principle for living structure. These methods show clearly the high explanatory power of deductive methods in morphology, but they also make one open end most explicit: neutral issues do exist. (shrink)
In this paper we reconsider the issue of free choice and the role of the whmorphology employed in it. We show that the property of being an interrogative whword alone is not sufficient for free choice, and that semantic and sometimes even morphological definiteness is a pre-requisite for some free choice items (FCIs) in certain languages, e.g. in Greek and Mandarin Chinese. We propose a theory that explains the polarity behaviour of FCIs cross-linguistically, and allows indefinite (Giannakidou 2001) as well (...) as definite-like FCIs. The difference is manifested as a lexical distinction in English between any (indefinite) and wh-ever (definite); in Greek it appears as a choice between a FCI nominal modifier (taking an NP argument), which illustrates the indefinite option, and a FC free relative illustrating the definite one. We provide a compositional analysis of Greek FCIs in both incarnations, and derive in a parallel manner the Chinese FCIs. Here the definite.. (shrink)
Morphological elements, or structures, are sorted into four categories depending on their level of anatomical isolation and the presence or absence of intrinsically identifying characteristics. These four categories are used to highlight the difficulties with the concept of structure and our ability to identify or define structures. The analysis is extended to the concept of homology through a discussion of the methodological and philosophical problems of the current concept of homology. It is argued that homology is fundamentally a similarity based (...) concept rather than a phylogenetic concept, and a proposal is put forth to return to a comparative context for homology. It is shown that for both the concepts of structure and homology ana priori assumption of stable underlying patterns (i.e. archetypes) is essential. (shrink)
In ancient Greek, the pitch accent of most words depends on the syllabification assigned to underlying representations, while a smaller, morphologically identifiable class of derived words is accented on the basis of the surface syllable structure, which results from certain contraction and deletion processes. Noyer 1997 proposes a cyclic analysis of these facts and argues that they are incompatible with parallel OT assumptions. His central claim is that the pre-surface syllabification to which accent is assigned in the bulk of (...) the Greek vocabulary does not occur at a “level privileged by UG,” such as the word level or the “cycle-final” level, but simply at an arbitrary point in the derivation. (p. 502). The implication is that extrinsic rule ordering is required to do justice to the accent system. Thus, Noyer’s work presents a challenge to any version of OT phonology. In this paper, I take up the challenge and argue that, although fully parallel OT may not be up to dealing with these accentual facts, the stratal version of OT based on Lexical Phonology and Phonology (stratal OT, or LPM-OT, Kiparsky 2000, to appear) provides a much better analysis of them than phonology with ordered rules does. (shrink)
The perennial fascination with the relationship between language and thought has generated much research across various disciplines. In recent years, commentators have called for closer examination of the connection between language acquisition and conceptual development (Bowerman & Levinson, 2001). Rather than assuming that language development always presupposes cognitive development, several researchers have started considering whether language learning could transform conceptual structure by making certain concepts available to the learner (e.g., de Villiers & Pyers, 1997; Gopnik & Choi, 1995; Bowerman, 1996).
In this study the relationship between functional morpholoy and evolutionary biology is analysed by confronting the main concepts in both disciplines.Rather than only discussing this connection theoretically, the analysis is carried out by introducing important practical and experimental studies, which use aspects from both disciplines. The mentioned investigations are methodologically analysed and the consequences for extensions of the relationship are worked out. It can be shown that both disciplines have a large domain of their own and also share a large (...) common ground. Many disagreements among evolutionary biologists can be reduced to differences in general philosophy (idealism vs. realism), selection of phenomenona (structure vs. function), definition of concepts (natural selection) and the position of the concept theory as an explaining factor (neutralists vs. selectionists, random variation, determinate selection, etc.). (shrink)
Clahsen has added to the body of evidence that, on average, regular and irregular inflected words behave differently. However, the dual-mechanism account he supports predicts a crisp distinction; the empirical data instead suggest a fuzzy one, more in line with single-mechanism connectionist models.
roots In the Lexicon of Malagasy we include an entry whose string part is vidy ('buy'). Its category is 'RT [AG, TH) ', indicating that it is a root and is associated with a two element set of theta roles, AGFNT and THEME. Semantically this entry is interpreted as a binary relation (= a two participant event), noted VIDY'.
In late winter of 1864, Charles Darwin received two folio volumes on radiolarians, a group of one-celled marine organisms that secreted siliceous skeletons of unusual geometry. The author, the young German biologist Ernst Haeckel (fig. 1), had himself drawn the figures for the extraordinary copper-etched illustrations that filled the second volume.1 The gothic beauty of the plates astonished Darwin (fig. 2 ), but he must also have been drawn to passages that applied his theory to construct the descent relations of (...) these little known creatures. He replied to Haeckel that the volumes "were the most magnificent works which I have ever seen, & I am proud to possess a copy from the author."2 Emboldened by his own initiative in contacting the famous naturalist, Haeckel, a few days later, sent Darwin a newspaper clipping that described a meeting of the Society of German Naturalists and Physicians at Stettin, which occurred during the previous autumn. The article gave an extended and laudatory account of Haeckel's lecture defending Darwin's theory.3 Darwin immediately replied in his second letter: "I.. (shrink)
Organisms are self-producing and self-maintaining, or autopoietic systems. Therefore, the course of evolution and adaptation of an organism is strongly determined by its own internal properties, whatever role external selection may play. The internal properties may either act as constraints that preclude certain changes or they open new pathways: the organism canalizes its own evolution. As an example the evolution of feeding mechanisms in salamanders, especially in the lungless salamanders of the family Plethodontidae, is discussed. In this family a large (...) variety of different feeding mechanisms is found. The authors reconstruct this evolutionary process as a series of bifurcation points of either constraints or opportunities forming a sequence of preconditions for the formation of a high-speed projectile tongue characteristic of tropical salamanders. Furthermore, it is shown how parallel evolution of seemingly unrelated domains within an organism such as respiratory physiology, life history biology and pattern of ontogeny has rather direct relevance to the feeding biology, thus demonstrating that organisms always evolve as wholes. (shrink)
We argue that although E-Z Reader does a good job in simulating many basic facts related to readers' eye movements, two phenomena appear to pose a challenge to the model. The first has to do with word length mediating the way compound words are identified; the second concerns the effects of initial fixation position in a word on eye behavior.
While the copula in Korean may attach to various consituents built around nominal hosts, it is not totally unconstrained as to the nature of its host. In particular, there are some post-nominal particles with which it cannot co-occur. Based on the classification of Yang (1972), and following much other work, Cho and Sells (1995) adopt the well-known template in (3) for the nominal system, with each position exemplified by (...) the particles shown in (1) and (2). (shrink)
A concept for a primitive angiospermous branch system is given in order to have a starting point for the derivation of the diverse and highly differentiated branch systems observed in contemporary angiosperms. Hitherto Troll's (1964, 1969) comparative study of the synflorescences in this plant group — developed out predominantly on herbaceous plants — was the most comprehensive and sophisticated treatment dealing with branch systems. Unfortunately, the work on tropical tree architecture by Hallé et al. (1978) has no reference to the (...) classical studies of Troll and his pupils. Thus Müller-Doblies and Weberling (1984) emphasized the high degree of terminological incompatibility between the two works. Angiosperms are seen as a monophyletic plant division. Consequently, the branch system of the first primitive angiosperms must be the starting point in the evolution of the abundant diversity of branch systems and growth forms of modern angiosperms. If it is accepted that primitive angiospermous shoots were terminated by a large flower, one may assume, that the reproductive end of the shoot was enriched by paracladia early in evolution, thereby developing a terminal inflorescence instead of a single flower. Thus the primitive shoot unit was divided into a basal vegetative region — the trophotagma, branching retardively — and the reproductive terminal region, the anthotagma, branching simultaneously. It is demonstrated through a selection of different examples, that the construction of such a system possesses the options for several modifications, enabling the evolution of the abundant diversity of branch systems which characterizes contemporary angiosperms. (shrink)
Organic selection (the Baldwin Effect) by which an environmentally elicitedphenotypic adaptation comes under genotypic control following selectionwas proposed independently in 1896 by the psychologists James Baldwinand Conwy Lloyd Morgan and by the paleontologist Henry Fairfield Osborn.Modified forms of organic selection were proposed as autonomization bySchmalhausen in 1938, as genetic assimilation by Waddington in 1942, andas an explanation for evolution in changing environments or for speciationby Matsuda and West-Eberhard in the 1980s. Organic selection as amechanism mediating proximate environmental effects on the (...) evolution ofmorphology and behaviour is the topic of this essay. Discussion includesthe context in which organic selection was proposed, Lamarckian or neo-Lamarckian implications of organic selection, Waddingtons experimentalstudies demonstrating the existence and efficacy of genetic assimilation,stabilizing selection and norms of reaction favoured by Schmalhausen, andMatsudas search for a mechanism of organic selection in endocrine changesand in heterochrony. (shrink)
The shapes of neurons and glial cells dictate many important aspects of their functions. In olfactory systems, certain architectural features are characteristics of these two cell types across a wide variety of species. The accumulated evidence suggests that these common features may play fundamental roles in olfactoryinformation processing. For instance, the primary olfactory neuropil in most vertebrate and invertebrate olfactory systems is organized into discrete modules called glomeruli. Inside each glomerulus, sensory axons and CNS neurons branch and synapse in patterns (...) that are repeated across species. In many species, moreover, the glomeruli are enveloped by a thin and ordered layer of glial processes. Theglomerular arrangement reflects the processing of odor information in modules that encode the discrete molecular attributes of odorant stimuli being processed. Recent studies of the mechanisms that guide the development of olfactory neurons and glial cells have revealed complex reciprocal interactions between these two cell types, which may be necessary for the establishment of modular compartments. Collectively, the findings reviewed here suggest that specialized cellular architecture plays key functional roles in the detection, analysis, and discrimination of odors at early steps in olfactory processing. (shrink)
The shapes of neurons and glial cells dictate many important aspects of their functions. In olfactory systems, certain architectural features are characteristics of these two cell types across a wide variety of species. The accumulated evidence suggests that these common features may play fundamental roles in olfactoryinformation processing. For instance, the primary olfactory neuropil in most vertebrate and invertebrate olfactory systems is organized into discrete modules called glomeruli. Inside each glomerulus, sensory axons and CNS neurons branch and synapse in patterns (...) that are repeated across species. In many species, moreover, the glomeruli are enveloped by a thin and ordered layer of glial processes. Theglomerular arrangement reflects the processing of odor information in modules that encode the discrete molecular attributes of odorant stimuli being processed. Recent studies of the mechanisms that guide the development of olfactory neurons and glial cells have revealed complex reciprocal interactions between these two cell types, which may be necessary for the establishment of modular compartments. Collectively, the findings reviewed here suggest that specialized cellular architecture plays key functional roles in the detection, analysis, and discrimination of odors at early steps in olfactory processing. (shrink)
The frequently occurring photoreceptor patterns in fish are explained using functional and environmental demands in a geometric model. The shape of the double cone provides a number of constructional properties leading to a limited number of appropriate configurations. The probability of their occurrence is estimated from the degree to which the combination of properties of each configuration meets specific environmental light conditions. A row pattern of merely double cones is especially suitable for vision in a dim homochromatic environment; a triangular (...) pattern is quite appropriate for high resolution and accurate movement detection, whereas the known square pattern has a high adaptive capacity to varying spectral distributions. In this context the transforming capacities of both square and row patterns can be understood. (shrink)
This book is concerned with the relationship between semantics and surface structure and in particular with the way in which each is mapped into the other. Jim Miller argues that semantic and syntactic structure require different representations and that semantic structure is far more complex than many analysts realise. He argues further that semantic structure should be based on notions of location and movement. The need for a semantic component of greater complexity is demonstrated by an examination of prepositions, particles, (...) adverbs and verb-prefixes, and is shown to accord with cross-language and historical facts. The volume goes on to consider the sort of rules that are required to map semantic structures onto syntax. Semantics and Syntax tackles fundamental issues and draws together many of the key concepts of traditional grammar and formal linguistics. The general framework for handling syntax, semantics and morphology that it outlines is perhaps a controversial one, but it will be recognized as challenging and original. (shrink)
This article deals with a type of functional explanation, viability explanation, that has been overlooked in recent philosophy of science. Viability explanations relate traits of organisms and their environments in terms of what an individual needs to survive and reproduce. I show that viability explanations are neither causal nor historical and that, therefore, they should be accounted for as a distinct type of explanation.
Chomsky and others have denied the relevance of external linguistic entities, such as E-languages, to linguistic explanation, and have questioned their coherence altogether. I discuss a new approach to understanding the nature of linguistic entities, focusing in particular on making sense of the varieties of kinds of “words” that are employed in linguistic theorizing. This treatment of linguistic entities in general is applied to constructing an understanding of external linguistic entities.
In a first part I present the results of the philosophy of scientific explanation with an attempt to apply them to the case of the theory of evolution. Then I observe that the requirements of modelization of phenomena with the help of inductive logic do not capture efficiently the pertinent factors and fail just as much to exclude those which end up being neutral as explanatory premises. I then query in the direction of confirmation theory, and show that probabilistic reasoning (...) does not possess the syntactic means of testing evolutionary hypotheses in a way that would rely on a valid mode of inference. In a second part, I try to show how biology has oscillated between two privileged modes of explanation, the first one being through form of which I suggest here that it has never been replaced and that it has known a forgotten vitality in the middle and latter part of the 19th Century, and the second one being through function here evaluated critically alongside that of tinkering. Finally, some limits are seen to hold against the pretence to epistemological exclusivism of those two outlooks on the biological object. (shrink)
This paper reports on the investigation of the effects of neuronal shape, at both individual cell and network level, on the behavior of neuronal systems. More specifically, two-dimensional biologically realistic neuronal networks are obtained that take explicity into account the position and morphology of neuronal cells, with the respective behavior for associative recall being simulated through a diluted version of Hopfield's model. While a specific probability density function is used for the placement of the cell bodies, images of real (...) neuronal cells (namely alpha and beta ganglion cells from the cat retina) are used to obtain biologically realistic models. Several morphological measures – including fractal dimension, the excluded volume, and integral geometry functionals–are estimated for the considered cells, and their values are correlated with the potential of the network for associative recall, which is quantified in terms of the overlap between distorted version of the trained patterns and their original version. Such an approach allows the quantitative and objective characterization of the relationship between neuronal shape and function, an important issue in neuroscience. The obtained results substantiate interesting relationships between the neural morphology and function as determined by the performance of the network. (shrink)
When phylogenetic trees constructed from morphological and molecular evidence disagree (i.e. are incongruent) it has been suggested that the differences are spurious or that the molecular results should be preferred a priori. Comparing trees can increase confidence (congruence), or demonstrate that at least one tree is incorrect (incongruence). Statistical analyses of 181 molecular and 49 morphological trees shows that incongruence is greater between than within the morphological and molecular partitions, and this difference is significant for the molecular partition. Because the (...) level of incongruence between a pair of trees gives a minimum bound on how much error is present in the two trees, our results indicate that the level of error may be underestimated by congruence within partitions. Thus comparisons between morphological and molecular trees are particularly useful for detecting this incongruence (spurious or otherwise). Molecular trees have higher average congruence than morphological trees, but the difference is not significant, and both within- and between-partition incongruence is much lower than expected by chance alone. Our results suggest that both molecular and morphological trees are, in general, useful approximations of a common underlying phylogeny and thus, when molecules and morphology clash, molecular phylogenies should not be considered more reliable a priori. (shrink)
The distinction between regular and irregular morphology is not clear-cut enough to suggest two distinct modular structures. Instead, regularity is tied directly to the type frequency of a pattern. Evidence from experiments as well as from naturally occurring sound change suggests that even regular forms have lexical storage. Finally, the development trajectory entailed by the dual-processing model is much more complex than that entailed by associative network models.
Dicotyledons are polarly organised in several ways. In plant morphology polarity, a principle allowing comparison of different plant structures has until yet not been studied. A division** of the vegetative plant in shoot and root as polar structures leads to the distinction of four instead of three basic organs: leaf, shoot axis, root axis and root cap together with the root hairs. The flower is also polarly organised, its poles are formed by the carpels and the stamens. The foliage (...) leaves are also polarly organised which is reflected by the morphological relationship of foliage leaf, stamen and carpel. The stamen uses the hypophyll*** as base of construction and the carpel uses the epiphyll**** as base of construction. Hypophyll and epiphyll are the two poles of the foliage leaf. Root and shoot, the polar entities of the vegetative plant and stamen and carpel, the polar entities of the generative plant, are morphologically correlated. Stamen and carpel can be understood as a combination of the basic organs of vegetative and generative parts of the plant. The basic organs of the generative plant are pollen grain and embryo sack with their gametophytes. The quantitative comparison of variable proportions is supplemented by a qualitative comparison of polarities. The result is, that the organisation type of the dicotyledons can yet be understood as constituted of morphological related parts.* in german Grundorgane. (shrink)
A paradox at the heart of language acquisition research is that, to achieve adult-like competence, children must acquire the ability to generalize verbs into non-attested structures, while avoiding utterances that are deemed ungrammatical by native speakers. For example, children must learn that, to denote the reversal of an action, un- can be added to many verbs, but not all (e.g., roll/unroll; close/*unclose). This study compared theoretical accounts of how this is done. Children aged 5–6 (N = 18), 9–10 (N = (...) 18), and adults (N = 18) rated the acceptability of un- prefixed forms of 48 verbs (and, as a control, bare forms). Across verbs, a negative correlation was observed between the acceptability of ungrammatical un- prefixed forms (e.g., *unclose) and the frequency of (a) the bare form and (b) alternative forms (e.g., open), supporting the entrenchment and pre-emption hypotheses, respectively. Independent ratings of the extent to which verbs instantiate the semantic properties characteristic of a hypothesized semantic cryptotype for un- prefixation were a significant positive predictor of acceptability, for all age groups. The relative importance of each factor differed for attested and unattested un- forms and also varied with age. The findings are interpreted in the context of a new hybrid account designed to incorporate the three factors of entrenchment, pre-emption, and verb semantics. (shrink)
In his article Forme et Fonction Barge wrote in 1936 that living matter cannot be totally understood in terms of causality. In this paper we argue on the contrary that this is in principle possible.In order to develop our arguments, we discuss some basic and derived concepts used in morphology and functional anatomy. We also formulate comments on the so-called formal, functional and final elucidations.
Cognition or higher brain activity is sometimes seen as a phenomenon greater than the sum of its parts. This viewpoint however is largely dependent on the state of the art of experimental techniques that endeavor to characterize morphology and its association to function. Retinal ganglion cells are readily accessible for this work and we discuss recent advances in computational techniques in identifying novel parameters that describe structural attributes possibly associated with specific function. These parameters are based on calculating wavelet (...) gradients from cell images followed by the extraction of meaningful measures including 2nd wavelet moment, entropy of orientation, and curvature. For the three cell types analyzed, the mean 2nd wavelet moment, which relates to the field of influence of the dendritic-tree segments was significantly different. cells had the highest mean 2nd wavelet moment, followed by the and cells (134 ± 22, 93 ± 19 and 63 ± 12, respectively). There was no significant difference between cells for entropy of orientation, indicating no class with a preferential orientation of their dendritic tree. Curvature provided similar results to the 2nd wavelet moment with cells having the highest curvature followed by and the cells (mean ± SD: 161 ± 15; 134 ± 22; 121 ± 15). Our feature space analysis also indicated a difference between these cell types. No difference was found between the and cell types and their physiological counterparts the Y and X cells based on wavelet analysis. Both the X and Y cells can be divided into two subtypes, the ON- and OFF-center cells based on the stratification level of the dendritic tree within the retina. Using 2nd wavelet moment, a difference in their morphological attributes, not reported previously, was noted for these subtypes. The 2nd wavelet moment and curvature are further discussed with respect to explaining regularity of spacing and coverage associated with retinal ganglion cell mosaics. (shrink)
This paper reports on the investigation of the effects of neuronal shape, at both individual cell and network level, on the behavior of neuronal systems. More specifically, two-dimensional biologically realistic neuronal networks are obtained that take explicity into account the position and morphology of neuronal cells, with the respective behavior for associative recall being simulated through a diluted version of Hopfield's model. While a specific probability density function is used for the placement of the cell bodies, images of real (...) neuronal cells (namely alpha and beta ganglion cells from the cat retina) are used to obtain biologically realistic models. Several morphological measures â including fractal dimension, the excluded volume, and integral geometry functionalsâare estimated for the considered cells, and their values are correlated with the potential of the network for associative recall, which is quantified in terms of the overlap between distorted version of the trained patterns and their original version. Such an approach allows the quantitative and objective characterization of the relationship between neuronal shape and function, an important issue in neuroscience. The obtained results substantiate interesting relationships between the neural morphology and function as determined by the performance of the network. (shrink)
According to rationalism regarding the psychology of moral judgment, people’s moral judgments are generally the result of a process of reasoning that relies on moral principles or rules. By contrast, intuitionist models of moral judgment hold that people generally come to have moral judgments about particular cases on the basis of gut-level, emotion-driven intuition, and do so without reliance on reasoning and hence without reliance on moral principles. In recent years the intuitionist model has been forcefully defended by Jonathan Haidt. (...) One important implication of Haidt’s model is that in giving reasons for their moral judgments people tend to confabulate – the reasons they give in attempting to explain their moral judgments are not really operative in producing those judgments. Moral reason-giving on Haidt’s view is generally a matter of post hoc confabulation. Against Haidt, we argue for a version of rationalism that we call ‘morphological rationalism.’ We label our version ‘morphological’ because according to it, the information contained in moral principles is embodied in the standing structure of a typical individual’s cognitive system, and this morphologically embodied information plays a causal role in the generation of particular moral judgments. The manner in which the principles play this role is via ‘proceduralization’ – such principles operate automatically. In contrast to Haidt’s intuitionism, then, our view does not imply that people’s moral reason-giving practices are matters of confabulation. In defense of our view, we appeal to what we call the ‘nonjarring’ character of the phenomenology of making moral judgments and of giving reasons for those judgments. (shrink)
Morphological content is information that is implicitly embodied in the standing structure of a cognitive system and is automatically accommodated during cognitive processing without first becoming explicit in consciousness. We maintain that much belief-formation in human cognition is essentially morphological : i.e., it draws heavily on large amounts of morphological content, and must do so in order to tractably accommodate the holistic evidential relevance of background information possessed by the cognitive agent. We also advocate a form of experiential evidentialism concerning (...) epistemic justification—roughly, the view that the justification-status of an agent’s beliefs is fully determined by the character of the agent’s conscious experience. We have previously defended both the thesis that much belief-formation is essentially morphological, and also a version of evidentialism. Here we explain how experiential evidentialism can be smoothly and plausibly combined with the thesis that much of the cognitive processing that generates justified beliefs is essentially morphological. The leading idea is this: even though epistemically relevant morphological content does not become explicit in consciousness during the process of belief-generation, nevertheless such content does affect the overall character of conscious experience in an epistemically significant way: it is implicit in conscious experience, and is implicitly appreciated by the experiencing agent. (shrink)
In the formation of epistemically justified beliefs, what is the role of attention, and what is the role (if any) of non-attentional aspects of cognition? We will here argue that there is an essential role for certain nonattentional aspects. These involve epistemically relevant background information that is implicit in the standing structure of an epistemic agent’s cognitive architecture and that does not get explicitly represented during belief-forming cognitive processing. Since such “morphological content” (as we call it) does not become explicit (...) during belief formation, it cannot be information that is within the scope of attention. Nevertheless,it does exert a subtle influence on the character of conscious experience, rather than operating in a purely unconscious way. (shrink)
After a historical sketch of the dynamical hypothesis, we stress that it is a functionalist hypothesis. We then tackle the point of a dynamical approach to constituent structures and emphasize that dynamical modeling must be coupled with morphological analysis.
Clahsen's conception of inflectional rules – as being not only regular, but simultaneously only concatenative (combinatorial), general and productive, representing the default, not occurring in interfixation within German compounds, and identical to the first rules to be acquired in first-language acquisition – involves an unwarranted collapsing of morphological concepts.
A method of phylogenetic reconstruction as proposed by a number of scientists of the Senckenberg Research Institute is discussed. The method is based on functional-morphological studies, the evolutionary adaptation principle of Bock and Von Wahlert (1965) and so-called model reconstruction. It is argued in this paper that direction of the adaptation process cannot be determined because of lack of knowledge about particular selective forces and that theories of model reconstruction are not open to contradiction in the sense of Popperian falsification. (...) Although it has been claimed that the method provides the only valid directional argument for morphoclines in cladistic studies, it remains unclear how to proceed when morphoclines show contradictory polarities. Moreover, it is doubtful whether polarities of morphoclines can be determined independently of phylogenetic hypotheses, and also whether the use of multistate morphoclines is methodologically valid. By relying on a particular evolutionary theory, i.e. the neo-Darwinian theory, and consequently assigning natural selection as the major agent of directional progress, the Senckenburg method of phylogenetic reconstruction restricts itself to microevolutionary change and, therefore, cannot be used when other hypotheses on the evolutionary process appear to explain the speciation process more plausibly, i.e. hypotheses on macroevolution. Furthermore, it is an unproved statement that evolution always proceeds according to the principle of economy. (shrink)
The article defends the doctrine that Linnaean taxa, including species, have essences that are, at least partly, underlying intrinsic, mostly genetic, properties. The consensus among philosophers of biology is that such essentialism is deeply wrong, indeed incompatible with Darwinism. I argue that biological generalizations about the morphology, physiology, and behavior of species require structural explanations that must advert to these essential properties. The objection that, according to current “species concepts,” species are relational is rejected. These concepts are primarily concerned (...) with what it is for a kind to be a species and throw little light on the essentialist issue of what it is for an organism to be a member of a particular kind. Finally, the article argues that this essentialism can accommodate features of Darwinism associated with variation and change. *Received August 2007; revised May 2008. †To contact the author, please write to: Philosophy Program, Graduate Center of the City University of New York, 365 Fifth Avenue, New York, NY 10016; e-mail: mdevitt@gc.cuny.edu . Essentialism about species is today a dead issue. (Sober [1980] 1992 , 249) Folk essentialism is both false and fundamentally inconsistent with the Darwinian view of species. (Griffiths 2002 , 72). (shrink)
Cognitive systems research has predominantly been guided by the historical distinction between emotion and cognition, and has focused its efforts on modelling the “cognitive” aspects of behaviour. While this initially meant modelling only the control system of cognitive creatures, with the advent of “embodied” cognitive science this expanded to also modelling the interactions between the control system and the external environment. What did not seem to change with this embodiment revolution, however, was the attitude towards affect and emotion in cognitive (...) science. This paper argues that cognitive systems research is now beginning to integrate these aspects of natural cognitive systems into cognitive science proper, not in virtue of traditional “embodied cognitive science”, which focuses predominantly on the body’s gross morphology, but rather in virtue of research into the interoceptive, organismic basis of natural cognitive systems. (shrink)
This paper raises fundamental questions about the claims of art historian David Freedberg and neuroscientist Vittorio Gallese in their article "Motion, Emotion and Empathy in Esthetic Experience." It does so from several perspectives, all of them rooted in the dynamic realities of movement. It shows on the basis of neuroscientific research how connectivity and pruning are of unmistakable import in the interneuronal dynamic patternings in the human brain from birth onward. In effect, it shows that mirror neurons are contingent on (...)morphology and corporeal-kinetic tactile-kinesthetic experience. Accordingly, it poses and answers the overlooked but seminally important question of how mirror neurons come to be. The original neuromuscular research of Parma neuroscientists and the findings of Marc Jeannerod concerning kinesthesia support the answer that the "underpinnings" of visual art appreciation are themselves underpinned. An abbreviated phenomenological analysis of movement and its implications regarding the fact that the making of all art is quintessentially contingent on movement, hence a dynamic enterprise, further bolster the given answer as does a brief review of an empirical phenomenological analysis of the natural dynamic congruency of emotions and movement. In the end, the paper shows that movement and life are of a piece in the creation and appreciation of art as in everyday life. (shrink)
Contemporary essentialism and real essentialism -- Against modalism -- Reductionism : the illusory search for inner constitution -- Why real essentialism? -- Some varieties of anti-essentialism -- Empiricist anti-essentialism -- Quinean animadversions -- Popper : avoiding what-is questions -- Wittgenstein : the shadow of grammar -- The reality and knowability of essence -- Why essences are real -- The problem of the universal accidental -- An empirical test for essence? -- Coming to know essence -- Paradigms, stereotypes, and classification -- (...) The structure of essence -- Hylemorphism : act and potency -- Substantial form -- Prime matter -- Substance -- The immanence of essence -- Essence and identity -- Real definition and the true law of identity -- The porphyrian tree -- The analogy of being -- Individuation -- Identity over time -- Essence and existence -- The real distinction in contingent beings -- Everything is contingentalmost -- Powers -- Laws of nature -- Aspects of essence -- Kinds of accident -- The nature of properties -- Knowledge of essence via properties -- Artefacts -- Origin and constitution -- Life -- The essence of life -- Kinds of organism -- Against emergence -- Species, biological and metaphysical -- Is biological essentialism dead? -- Against the cladistic species concept -- Vagueness -- A plea for morphology -- The person -- The essence of personhood -- Hylemorphic dualism -- Consciousness, psychology, and the person -- Form, body, and soul -- Soul, intellect, and immateriality -- Soul, identity, and material dependence. (shrink)
We examine the question of which aspects of language are uniquely human and uniquely linguistic in light of recent suggestions by Hauser, Chomsky, and Fitch that the only such aspect is syntactic recursion, the rest of language being either specific to humans but not to language (e.g. words and concepts) or not specific to humans (e.g. speech perception). We find the hypothesis problematic. It ignores the many aspects of grammar that are not recursive, such as phonology, morphology, case, agreement, (...) and many properties of words. It is inconsistent with the anatomy and neural control of the human vocal tract. And it is weakened by experiments suggesting that speech perception cannot be reduced to primate audition, that word learning cannot be reduced to fact learning, and that at least one gene involved in speech and language was evolutionarily selected in the human lineage but is not specific to recursion. The recursion-only claim, we suggest, is motivated by Chomsky’s recent approach to syntax, the Minimalist Program, which de-emphasizes the same aspects of language. The approach, however, is sufficiently problematic that it cannot be used to support claims about evolution. We contest related arguments that language is not an adaptation, namely that it is “perfect,” non-redundant, unusable in any partial form, and badly designed for.. (shrink)
A series of representations must be semantics-driven if the members of that series are to combine into a single thought. Where semantics is not operative, there is at most a series of disjoint representations that add up to nothing true or false, and therefore do not constitute a thought at all. There is necessarily a gulf between simulating thought, on the one hand, and actually thinking, on the other. A related point is that a popular doctrine - the so-called 'computational (...) theory of mind' (CTM) - is based on a confusion. CTM is the view that thought-processes consist in 'computations', where a computation is defined as a 'form-driven' operation on symbols. The expression 'form-driven operation' is ambiguous, and may refer either to syntax-driven operations or to morphology-driven operations. Syntax-driven operations presuppose the existence of operations that are driven by semantic and extra-semantic knowledge. So CTM is false if the terms 'computation' and 'form-driven operation' are taken to refer to syntax-driven operations. Thus, if CTM is to work, those expressions must be taken to refer to morphology-driven operations; and CTM therefore fails, given that an operation must be semantics-driven if it is to qualify as a thought. CTM therefore fails on every disambiguation of the expressions 'formal operation' and 'computation,' and it is therefore false. (shrink)
