Aristotle: Biology

Vortrag, gehalten am 11. November 2020 im Rahmen einer Ringvorlesung am Regensburger Zentrum für Klassikstudien zum Thema "Entfernte Verwandte - Mensch und Tier". Die aristotelische Bestimmung des Menschen ist ein Rätsel. Daher soll sie im Folgenden auch als ein Rätsel behandelt werden. Ziel ist es, hier nicht das bei Aristoteles finden zu wollen, was wir heute ohnehin schon über den Menschen als ein Lebewesen wissen oder zu wissen glauben, sondern es gilt im Folgenden von Aristoteles ahnen zu lernen, was wir (...) noch nicht wissen, wofür wir heute, in unserer technisierten, wissenschaftsorientierten und digitalisierten Blickbahn vielleicht noch gar nicht gerüstet sind. (shrink)

It has become somewhat of a platitude to call Aristotle the first epigenesist insofar as he thought form and structure emerged gradually from an unorganized, amorphous embryo. But modern biology now recognizes two senses of “epigenesis”. The first is this more familiar idea about the gradual emergence of form and structure, which is traditionally opposed to the idea of preformationism. But modern biologists also use “epigenesis” to emphasize the context-dependency of the process itself. Used in this sense development is not (...) simply the unfolding of a pre-determined sequence of changes specified in advance by the organism’s genotype. It is also sensitive to inputs from the internal and external environment, which help determine in real-time which of the many potential developmental pathways are actualized during the process. Within this paradigm developing embryos are viewed as dynamic and responsive systems that react to inputs from the internal and external environment ‘on the fly’. In this paper I argue that, while Aristotle was an epigenesist in the first sense, he would have rejected epigenesis in the more modern sense. First, Aristotle’s model of choice for a developing embryo is the automaton that executes a set of preset movements (GA 734b9-13, 741b7-15). The automatons he has in mind are not dynamic AI systems capable of modifying their behaviour on the fly in response to environmental cues but completely deterministic mechanisms whose movements are fixed by their original design. Second, given Aristotle’s views about the different kinds of causal powers there are, it looks like only intentional agents endowed with actual decision-making powers could be capable of the sort of plasticity at the core of a more dynamic epigenesis. For that kind of epigenesis requires powers for alternative outcomes, and Aristotle is explicit that such powers require rational desires (προαίρεσις) that control which of those alternatives to bring about. If I am right, then he could not have made sense of the idea of a developing embryo (as a non-intentional system) making adjustments to its phenotype on the fly in response to emerging problems and opportunities, given the conceptual resources available to him. (shrink)

The intellectual history of evolutionary theory really does not begin in earnest until the late seventeenth/early eighteenth century. Prior to that, the idea that species might have evolved over time was not a serious possibility for most naturalists and philosophers. There is certainly no substantive debate in antiquity about evolution in the modern sense. There were really only two competing explanations for how living things came to have the parts they do: design or blind chance. Ancient Greek Atomism, for example, (...) taught that all composite bodies, including living things, are generated through the random collision of atoms as they rebel and move in the void. Plato and Aristotle both dismissed this possibility on the grounds that living things are too complex and too well-adapted to be products of chance. This eventually became the central premise in Galen’s own Argument from Design. That species forms might have gradually evolved over time by a process of natural selection was not seen as a plausible alternative. Of course, such a theory had been proposed by Empedocles in the fifth century BCE. But because his theory still relied heavily on chance, it was not taken seriously by any of the later ancient Greek or Roman thinkers. My aim in this paper is to investigate the reasons why evolutionary thinking failed to gain momentum in antiquity after its introduction by Empedocles. (shrink)

Aristotle's voluminous writings on animals have often been marginalised in the history of philosophy. Providing the first full-length comprehensive account of Aristotle's biology, its background, content and influence, this Companion situates his study of living nature within his broader philosophy and theology and differentiates it from other medical and philosophical theories. An overview of empiricism in Aristotle's Historia Animalium is followed by an account of the general methodology recommended in the Parts of Animals. An account of the importance of Aristotle's (...) teleological perspective and the fundamental metaphysics of biological entities provides a basis for understanding living capacities, such as nutrition, reproduction, perception and self-motion, in his philosophy. The importance of Aristotle's zoology to both his ethics and political philosophy is highlighted. The volume explores in detail the changing interpretations and influences of Aristotle's biological works from antiquity to modern philosophy of science. It is essential for both students and scholars. (shrink)

In De Anima 2.4, Aristotle claims that nutritive soul encompasses two distinct biological functions: nutrition and reproduction. We challenge a pervasive interpretation which posits ‘nutrients’ as the correlative object of the nutritive capacity. Instead, the shared object of nutrition and reproduction is that which is nourished and reproduced: the ensouled body, qua ensouled. Both functions aim at preserving this object, and thus at preserving the form, life, and being of the individual organism. In each case, we show how Aristotle’s detailed (...) biological analysis supports this ontological argument. (shrink)

The soul is an explanatory principle of Aristotle’s natural science, accounting both for the fact that living things are alive as well as for the diverse natural attributes that belong to them by virtue of being alive. I argue that the explanatory role of the soul in Aristotle’s natural science must be understood in light of his view, stated in a controversial passage from Parts of Animals (645b14–20), that the soul of a living thing is a “complex activity” of its (...) organic body. This paper explores the role of this “complex activity” model of soul in Aristotle’s study of soul in De Anima. I argue, first, that the model has its origins in De Anima II.4, where Aristotle argues that living things do all they do by nature for the sake of a single, teleologically primary end. I argue further that Aristotle uses this model to account for the psychological attributes naturally present in living things, including their capacities for vital activities like nutrition, reproduction, and perception, and that this is the task to which Aristotle devotes the obscure final chapters of De Anima III. (shrink)

Hynek Bartos does the field of ancient philosophy a great service by detailing the influence of early Greek thinkers (such as Heraclitus, Empedocles, Anaxagoras, Democritus, and Diogenes of Apollonia) on the Hippocratic work On Regimen, and by demonstrating that work’s innovative engagement with contemporary scientific and philosophical concepts as well as its direct influence on Plato and Aristotle. His study usefully counteracts the lamentable tendency among ancient philosophers to ignore or downplay the influence of medical literature on philosophy in general, (...) and to heap scorn on this text in particular, which Kirk, Raven, and Schofield considered ‘an eclectic and very superficial quasi- philosophical treatise’ and Barnes famously called ‘a silly farrago of ill-digested Presocratic opinions’. (shrink)

Der Aufsatz möchte, ausgehend von einer zureichenden Darstellung des thematischen Bereichs von Ernährung und Kühlung bei Aristoteles (Abschnitt 1.1) sowie seiner Physiologie der Geruchswahrnehmung im allgemeinen (Abschnitt 1.2), zum einen zeigen, wie Aristoteles das Gegebensein der Wahrnehmung von Düften beim Menschen zoologisch und physiologisch begründet und dabei medizinische Implikationen formuliert (Abschnitt 2); zum anderen wird zu sehen sein, inwiefern die Wahrnehmung von Düften ethische und ästhetische Züge aufweist, wofür das entsprechende zoologische und physiologische Wissen den ausdrücklichen oder unausdrücklichen Horizont zu (...) bilden vermag (Abschnitt 3). In diesem Sinne bietet das Beispiel der spezifischen Geruchswahrnehmung des Menschen die Möglichkeit, zu zeigen, inwiefern die Zoologie, im größeren Zusammenhang des aristotelischen Denkens, für die genannten Wissensfelder ein Grundlagenwissen eröffnen kann (Abschnitt 4). (shrink)

Aristotle contrasts standard animal generation with ‘spontaneous generation’, which happens when some material putrefies and gives rise to a new organism. This paper addresses two interrelated puzzles about spontaneous generation. First, is it of the same ‘fundamental kind’ of causal process as standard generation? Second, is it ‘spontaneous’, as understood in Physics 2.4–6: rare, accidentally caused, and among things that are for the sake of something? I argue that both puzzles turn on the same questions about the process types involved. (...) I show that the type putrefaction plays a more important role in spontaneous generation than has been recognized so far. Because putrefaction does not play this role in standard generation, the two processes are of different ‘fundamental’ kinds. Moreover, spontaneous generation happens rarely in that it is rare for processes of putrefaction to happen in such a way that they can also be described in terms of concoction. (shrink)

The paper addresses the following question: why do human beings, on Aristotle’s view, have an innate tendency to badness, that is, to developing desires that go beyond, and often against, their natural needs? Given Aristotle’s teleological assumptions (including the thesis that nature does nothing in vain), such tendency should not be present. I argue that the culprit is to be found in the workings of rationality. In particular, it is the presence of theoretical reason that necessitates the limitless nature of (...) human non-rational desires. (shrink)

In this work I propose a theory of evolution as a process of unfolding. This theory is based on four logically concatenated principles. The principle of evolutionary order establishes that the more complex cannot be generated from the simpler. The principle of origin establishes that there must be a maximum complexity that originates the others by logical deduction. Finally, the principle of unfolding and the principle of actualization guarantee the development of the evolutionary process from the simplest to the most (...) complex. These logical principles determine the existence of a virtual ideological matrix that contains the sequence of the preformed and folded morphogenetic fields. In this manner, the evolutionary process consists of the sequential unfolding and actualization of these fields, which is motorized by a process of teleologization carried out by the opening consciousness of the forms included in the fields of the ideological matrix. This theory leads to a radical change of perspective regarding the materialist worldview, and places life at the center of the evolutionary process as an activity carried out by a consciousness that seeks to fulfill a purpose by actualizing its own potentialities. (shrink)

Diogenes subscribes to a principle that, roughly, causal interaction and change require a certain sort of uniformity among the relata. Attending to this principle can help us understand Diogenes's relationship to the superficially similar Anaximenes without insisting, as some do, that Diogenes must be consciously responding to Parmenides. Diogenes is distinctive and philosophically interesting because his principle combines two senses of ‘archê’ (principle, starting-point), namely, the idea of source or origin and that of underlying (material) principle, and gives the rudiments (...) of an argument for associating the two, by which Aristotle may have been influenced. Diogenes’s principle and its deployment in biological explanations thematized a concern that Aristotle at least partially shared, and which led him to appeal, as Diogenes is said to have done, to pneuma (breath, air). (shrink)

Chance and necessity are mainstays of explanation in current biology, dominated by the neo-Darwinian outlook, a blend of the theory of evolution by natural selection with the basic tenets of population genetics. In such a framework the form of living organisms is somehow a side effect of highly contingent, historical accidents. Thus, at a difference of other sciences, biology apparently lacks theoretical principles that in a law-like fashion may explain the emergence and persistence of the characteristic forms of living organisms (...) that paradoxically, given the current importance attributed to chance, can be grouped into organized structural typologies. Nevertheless, the present essay shows that since its origins in Aristotelian natural history, biology aimed at achieving rational, non-accidental, explanations for the wide variety of living forms endowed with characteristic behaviors that constitute the landscape of biological species. (shrink)

What is a human body? Some reasons are given for thinking that, in the primary case, it is a body that is both of and suitable to a rational animal.

Com este artigo, pretendo examinar a maneira pela qual ocorreria a necessidade natural, em seus diversos aspectos, nos distintos processos gerativos composicionais em Aristóteles. Em um caso, a necessidade natural se daria de um modo "sem mais", ou de um modo absoluto, por meio da qual se geram os agregados. Em outro, a necessidade natural se realizaria a partir de um princípio anterior regulativo ou determinante, que Aristóteles denomina de necessidade ex hupoteseos, com relação aos processos envolvidos na constituição dos (...) corpos homogêneos inanimados e, dos organismos vivos. No entanto, haveria uma diferença essencial relativamente ao acabamento composicional associado, por um lado, aos corpos homogêneos inanimados e, por outro, aos organismos vivos. Enquanto que o acabamento constituinte das composições homogêneas inanimadas se restringiria apenas ao todo composicional e suas propriedades características, o acabamento dos organismos vivos corresponderia ao todo composicional e as suas propriedades características, bem como este acabamento em vista da realização das atividades orgânico-funcionais, ou vitais. (shrink)

There are two different notions of seed at work in the Generation of Animals: seed as the spermatic residue, which concerns only the male and the female generative contributions, and seed as the kuêma and first mixture of the two generative contributions. The latter is a notion of seed common to plants and animals. The passage in GA I.18, 724b12–22 where Aristotle distinguishes between these two notions of seed has been mistakenly discredited as inauthentic or simply as irrelevant for understanding (...) the seeds of animals. On the other hand, recent studies have rather focused on the seed as spermatic residue, paying little attention to the notion of seed as kuêma. In this paper I defend the authenticity and relevance of this passage and show how understanding the notion of seed as kuêma is essential to have a complete picture of Aristotle’s account of seed. This common notion of seed makes sense of Aristotle’s otherwise puzzling use of the word “sperma” to designate the seeds of plants, the kuêma, the fertilized eggs, and the first mixture of the two generative contributions. It also proves helpful in determining what the word ‘sperma’ stands for in key passages, such as Metaph. IX.7. (shrink)

In this article, I present in a systematic way Aristotle’s understanding of movement (kinêsis) as efficient cause in the Generation of Animals. This aspect of movement is not disclosed in the approach to movement as an incomplete activity in contrast to energeia, which has been extensively discussed in the literature. I explain in which sense movement is the efficient cause of generation and how this movement is related to the other factors, in particular the source of movement, the seminal fluid, (...) pneuma, and the vital heat, as well as to the form and the soul. The role of movement as efficient cause of generation shows how, for Aristotle, form and soul are inseparable from matter and movement in nature. Explaining the role of movement as efficient cause of generation also helps to make sense of the multiplicity of factors involved in the efficient causality of generation. The key to articulating these factors with one another is the identity of form of the generative movement, which is specifically the same in the source, in the tools, and in the substance that is coming into being. (shrink)

This is the first collection of essays devoted specifically to the nature and significance of Aristotle's anthropological philosophy, covering the full range of his ethical, metaphysical and biological works. The book is organised into four parts, two of which deal with the metaphysics and biology of human nature and two of which discuss the anthropological foundations and implications of Aristotle's ethico-political works. The essay topics range from human nature and morality to friendship and politics, including original discussion and fresh perspectives (...) on rationalism, the intellect, perception, virtue, the faculty of speech and the differences and similarities between human and non-human animals. Wide-ranging and innovative, the volume will be highly relevant for readers studying Aristotle as well as for anyone working on either ancient or contemporary philosophical anthropology. (shrink)

Given that they are chance events, there can be no scientific demonstration or knowledge of monsters. There are still, however, many recognizable elements of scientific explanation in Aristotle's Generation of Animals Book IV chapter 4. What happens in cases of monsters and deformities occurs in the process of generation, and there is much that we can know scientifically about this process—working from the animal’s essential attributes outward to factors that influence these processes. In particular, we find Aristotle looking for and (...) investigating whether “for the most part” correlations are causally relevant and linked to essence. Even though the birth of monsters is scientifically inexplicable, the features of animals that tend to produce them can be known about. I will explain how it is that producing more than one offspring per birth is a knowable feature of a kind. This feature, along with others, can then give us a much better grasp on what happens in particular instances of monstrous birth. (shrink)

From Natural Character to Moral Virtue in Aristotle, by LeunissenMariska. Oxford: Oxford University Press, 2017. Pp. vii + 216.

Was Aristotle the ‘father’ and founder of the epigenesis doctrine? Historically, I will argue, this question must be answered with ‘no’. Aristotle did not initiate and had no access to a debate that described itself in terms of ‘epigenesis’ and ‘preformation’, and thus cannot be considered the ‘father’ or founder of the epigenesis-preformation controversy in a literal sense. But many ancient accounts of reproduction and embryological development contain analogies to what early modern scientist called ‘epigenesis’ and ‘preformation’, and, in this (...) analogous sense, Aristotle can be considered a precursor of the epigenesis-preformation controversy. But is Aristotle’s position actually epigenetic, as most of the traditional interpreters hold, or preformationist, as some of the recent scholars believe? I will argue against the one-sidedness of both readings that Aristotle’s account of reproduction and heredity contains mainly epigenetic, but also a few preformationist characteristics. Whereas, for instance, Aristotle’s idea of a successive development of the embryo’s parts is doubtlessly epigenetic, Aristotle’s idea that the development of the embryo is an actualization and enlargement of potential parts, which are simultaneously present in the semen, can be considered a preformationist feature. (shrink)

This is an epistemologically-driven history of the concept of evolution. Starting from its inception, this work will follow the development of this pregnant concept. However, in contradistinction to previous attempts, the objective will not be the identification of the different meanings it adopted through history, but conversely, it will let the concept to be unfolded, to be explicated and to express its own inner potentialities. The underlying thesis of the present work is, therefore, that the path that leads to the (...) development of the concept of evolution is the path that studies the possibilities of the evolution of concepts, and that the historical reconstruction of its conceptual trajectory will shed light into potential and unexploited possibilities. This methodology will provide useful tools and resources for future developments of the concept. For example, it will define the concept of transmutation as a different conceptual trajectory deviating from the one corresponding to evolution, at the onset of the 19th century. Moreover, epigenesis will not be the opposing concept to evolution, but only to simultaneous and instantaneous generation. It will demonstrate that every important system of epigenesis drew upon some kind of formative power to explain development. More importantly, it will show that the problem of preformation cannot be overlooked, and that some kind of virtual preformation must be considered in order to address the problems of generation and development. (shrink)

We have reason to think that a fundamental goal of natural science, on Aristotle’s view, is to discover the essence-specifying definitions of natural kinds—with biological species as perhaps the most obvious case. However, we have in the end precious little evidence regarding what an Aristotelian definition of the form of a natural kind would look like, and so Aristotle’s view remains especially obscure precisely where it seems to be most applicable. I argue that if we can get a better understanding (...) of how the forms of natural kinds are or come to be known, and how they make things intelligible, we can get a better appreciation of the nature of form in general, as well as solve certain puzzles about form and definition. (shrink)

Aristotle’s theory of spontaneous generation offers many puzzles to those who wish to understand his theory both within the context of his biology and within the context of his more general philosophy of nature. In this paper, I approach the difficult and vague elements of Aristotle’s account of spontaneous generation not as weaknesses, but as opportunities for an interesting glimpse into the thought of an early scientist struggling to reconcile evidence and theory. The paper has two goals: to give as (...) charitable and full an account as possible of what Aristotle’s theory of spontaneous generation was, and to examine some of its consequences; and to reflect on Aristotle as a scientist, and what his comments reveal about how he approached a difficult problem. In particular, I propose that the well-recognized problem of the incompatibility between Aristotle’s concept of spontaneity and his theory of spontaneous generation presents an opportunity for insight into his scientific methodology when approaching ill-understood phenomena. (shrink)

This paper aims to undertake an examination of the exposition made in the aristotelian treatise De Incessu Animalium about the directional orientations (above and below, right and left, front and behind) mostly from its extra, inter and intratextual context. These directional orientations seem to be directly related to the principles of movement, perception and growing. The treatise starts from the points of origin (archai) of these functions in the animals and, from these functions, distinguish them. Therefore, functionality is established as (...) the first value, because it is the prism through which Aristotle perceives and think the right, above and front in the animals. On that account, it is corroborated the great importance of functionality in the aristotelian philosophy. Afterwards, it will be searched if (or how) the primacy of the trinominal formed by right, above and front is related to aspects of the greek imaginary contemporaneous with the given text. (shrink)

Aristotle’s principle that “nature does nothing in vain” (NDNIV) is central to his teleological approach to understanding organisms. First, we argue that James G. Lennox’s influential account of NDNIV is unsuccessful. Second, we propose an alternative account that includes a natural state model. According to a natural state model of development, an organism will develop toward its natural state unless interfering forces prevent that from happening. Third, we argue that this account also fits Aristotle’s discussion in the Generation of Animals (...) and can explain monstrosities in nature that at first sight seem to be counterexamples to NDNIV. Fourth, we take a broader look at NDNIV, arguing that it does not entail that all structures are teleological (since it accommodates the thought that by-products and some neglected items that Aristotle calls “tokens” are nonteleological), that it does not entail that there is only one function per organ, that NDNIV is not an anthropocentric principle, and that, perhaps surprisingly, the teleological structures of an organism can benefit others apart from the organism and its progeny. Finally, we argue that Aristotle endorsed an empirical justification of NDNIV wherein the principle is justified by its many successful applications. (shrink)

In some influential histories of ancient philosophy, teleological explanation and mechanistic explanation are assumed to be directly opposed and mutually exclusive alternatives. I contend that this assumption is deeply flawed, and distorts our understanding both of teleological and mechanistic explanation, and of the history of mechanistic philosophy. To prove this point, I shall provide an overview of the first systematic treatise on mechanics, the short and neglected work Mechanical Problems, written either by Aristotle or by a very early member of (...) his school. I will argue that the work is thoroughly Aristotelian in methodology, and that taking it seriously can deepen our understanding of Aristotle’s discussion of animal and human self-motion in the Physics and On the Movement of Animals. (shrink)

Human beings, according to Aristotle, are not the only political animals. Bees, wasps, ants and cranes are the other political species mentioned by Aristotle in the History of Animals. Politics, I, 2 confirms this point and makes the additional statement that human beings, if not the only political animals, are nevertheless more political than the other political animals. There has been a traditional scholarly agreement that the capacity for rational speech is the reason why human beings are more political. This (...) traditional view seems to have some support from the same chapter of Politics, where Aristotle addresses, right after having stated that human beings are more political, the role of language in human beings’ political life. This paper contests this traditional view about language and claims that it is the more political character of human beings which explains the political role of language and not vice versa. -/- . (shrink)

Organisms receded from view in much of twentieth-century biology, only to undergo a sort of renaissance at the start of the twenty-first. The story of why this should be so is complicated and fascinating, but belongs primarily to the history of biology. On the other hand, to the extent that it is so, a question naturally arises: what, after all, are organisms? This question has a long and complicated history of its own, both within and without of biology; an investigation (...) of this history yields some guidance as to how organisms might yet be conceived today. One suggestion borne of these investigations is this: organisms are, for better or worse, normatively delineated unities. (shrink)

Wie an Politik I 2 in Verbindung mit anderen Passagen aus dem Corpus Aristotelicum, v.a. aus seinen zoologischen Schriften, gezeigt werden kann, ist die besondere Fähigkeit des Menschen, sich mitzuteilen, nicht ohne seine spezifische Zoologie denkbar. Ebensowenig ist daher die besondere menschliche Art, Haus- und Staatswesen zu bilden, ohne seine zoologischen Besonderheiten vorstellbar. Die menschliche Fähigkeit, sich mitteilen zu können, weist so in seine spezifische Art des Mitseins und eröffnet dadurch das, was er mitzuteilen vermag, z.B. Recht und Unrecht. Im (...) Unterschied zur Interpretation bei Thomas, der in der Art des Mitteilbaren das spezifische Mitsein des Menschen gegründet sieht, deutet sich dergestalt umgekehrt das Mitsein des Menschen sowie die Art seiner Teilhabe am Dasein überhaupt als Grundzug seiner Mitteilung und des durch sie Mitteilbaren an. Der Beitrag enthält im Anhang einige für die Erörterungen wichtige Begriffsbestimmungen, so u.a. zu den aristotelischen Begriffen 'autarkeia' und 'energeia/entelecheia'. (shrink)

Diversität ist ein zentraler Topos einer Vielzahl aktueller gesellschaftlicher Diskurse. Was genau ist aber gemeint, wenn von „Diversität“ oder auch „Vielheit“ und „Vielfalt“ die Rede ist? Handelt es sich um eine beschreibende Kategorie oder aber um eine, die normativ konnotiert ist? Der vorliegende Band analysiert – im Hinblick auf die Debatten um Biodiversität –, welche Bedeutungen und Funktionen „Diversität“, „Vielheit“ und „Vielfalt“ als Kategorie, Befund und Norm zukommen. Dazu rücken die Buchbeiträge die komplexen begriffs- und ideengeschichtlichen Hintergründe dieser Konzepte in (...) den Blick, indem sie die für die Thematik zentralen philosophischen Programme von Aristoteles, Ockham, Kant, Leibniz, Bergson und Whitehead jeweils im Zusammenhang mit möglichen biologischen Bezugsebenen betrachten. Die leitende Annahme für diesen Ansatz ist: Die heutige Verwendung von „Biodiversität“ als eine deskriptive und präskriptive Aspekte umfassende Kategorie lassen sich nur angemessen würdigen, wenn man sich die kulturellen und historischen Kontexte der Konzeptionierung und Bewertung von Diversität, Vielheit und Vielfalt vergegenwärtigt. (shrink)

Is life a simple result of a conjunction of physico-chemical processes? Can be reduced to a mere juxtaposition of spatially determined events? What epistemology or world-view allows us to comprehend it? Aristotle built a novel philosophical system in which nature is a dynamical totality which is in constant movement. Life is a manifestation of it, and is formed and governed by the psyche. Psyche is the organizational principle of the different biological levels: nutritive, perceptive and intelective. Driesch's crucial experiment provided (...) empirical proof of the principle of life, which he called entelechy. Entelechy is an intensive manifoldness and cannot be comprehended by the usual extensive parameters. The entelechian's own ambiance is duration. This allows the reintroduction of the concept of teleology in the sphere of the living, understood not as a final cause, but as an order born from desire and leading to action. (shrink)

Exceptions are often cited as a counterargument against formal causation. Against this I argue that Aristotle explicitly allows for exceptions to essences in his biological writings, and that he has a means of explaining them through formal causation – though this means that he has to slightly elaborate on his general case theory from the Posterior Analytics, by supplementing it with a special case application in the biological writings. Specifically for Aristotle an essential predication need not be a universal predication. (...) Rather an essential predication is where a property belongs essentially to a species or genus. This essential predication serves a causal and explanatory role, and is not dependent on a corresponding universal predication. (shrink)