Abstract
The term biosocial evolution refers to mutual relations between different organisms and specially to functional systems composed of individuals belonging to a species , or to two or more different species .The main features of the biosocial evolutioni.e. historical development of the functional systems of biosocial order are: 1. functional differentiation of the individual components of corresponding systems, 2. coordination of the differentiated functions and as a result, 3. the intergration of this systems as functional wholes.The tropical rain forest is to be considered as one of the most advanced terrestrial ecosystems, caracterized by its extreme differentiation, high level of its integration and violent reactions to main forces.The functional differentiation of the components of ecosystems is considered as a manifestation of the development of their individual traits as members of this system.The mechanisms of both phylogenetical and biosocial evolution as well as some other questions related to the problem are briefly discussed in the text