Abstract
Scribal copying is investigated as a test case for the memetic and epidemiological models for explaining the distribution of cultural items. We may hypothesize that the incidence of errors could be low enough to allow two conditions for neo-Darwinian explanation (or an analogue of it) to be fulfilled: first, that there be a rather reliable mechanism for heredity, and second that occasional mutations might produce a version more likely to survive and be propagated than the exemplar. Scriptorial conventions are reviewed. Textual criticism is investigated. Finally, some attention is given to the psychology of language perception and production. In conclusion, it is argued that the memetic model for cultural transmission is not generally fecund while the epidemiological model is.