True ecotourism requires us to regain an understanding of the inextricable links between the habitats of a region, including its inhabitants, and their habits. With this systemic approach that integrates economic, ecological, and ethical dimensions, we define ecotourism as “an invitation to a journey (‘tour’) to appreciate and share the ‘homes’ (oikos) of diverse human and non-human inhabitants, their singular habits and habitats.” Today, mass nature tourism often denies theselinks and is generating biocultural homogenization, socio-ecological degradation, and marked distributive injustices (...) in iconic places, such as Costa Rica, the Galapagos and Cape Horn. In order to implement genuine ecotourism in Latin America and elsewhere, it is imperative to overcome marketing ambiguities, and pay close attention to local autonomy and biocultural diversity. (shrink)

"Evolutionary ecologist and photographer John Hess presents the Galâapagos in stunning photographs and insightful prose, celebrating the archipelago as a unique place to appreciate the achievements of Charles Darwin and other biologists as ...

In my book Posthuman Life: Philosophy at the Edge of the Human (Routledge 2014) I set out a recursive account of the conditions for posthumanity: the Disconnection Thesis (DT). The DT states that a being is posthuman iff: -/- 1) It has ceased to belong to WH (the "Wide Human" socio-technical network) as a result of technical alteration; 2) Or it is a wide descendent of such a being (outside WH) (PHL 112) -/- Jon Cogburn's "Galapagos objection" attempts to show (...) that there could be life forms whose evolution satisfies the disconnection criteria but which lack the degree of autonomy that elicits our moral concern with the prospect posthumans. In this post I respond to Jon's objection and briefly discuss other philosophical objections to my account of posthumanity. -/- Roden, David. 2012. “The Disconnection Thesis”. In The Singularity Hypothesis: A Scientific and Philosophical Assessment, A. Eden, J. Søraker, J. Moor & E. Steinhart (eds), 281–98. London: Springer. -/- Roden, David. 2014. Posthuman Life: Philosophy at the Edge of the Human. London: Routledge. -/- . (shrink)

The target article excludes ancestral neutrality as a cause for the inheritance of schizophrenia, with an argument relating to selection against a single allele in the Finnish population. However, drift would predominate over selection within subisolates of the Finnish population. Comparisons of subisolates with heterogeneous populations may provide clues to the endophenotypic structure of complex polygenetic heritable mental disorders. (Published Online November 9 2006).

During his historic Galápagos visit in 1835, Darwin spent nine days making scientific observations and collecting specimens on Santiago (James Island). In the course of this visit, Darwin ascended twice to the Santiago highlands. There, near springs located close to the island's summit, he conducted his most detailed observations of Galapagos tortoises. The precise location of these springs, which has not previously been established, is here identified using Darwin's own writings, satellite maps, and GPS technology. Photographic evidence from excursions (...) to the areas where Darwin climbed, including repeat photography over a period of four decades, offers striking evidence of the deleterious impact of feral mammals introduced after Darwin's visit. Exploring the impact that Darwin's Santiago visit had on his thinking - especially focusing on his activities in the highlands - raises intriguing questions about the depth of his understanding of the evolutionary evidence he encountered while in the Galápagos. These questions and related insights provide further evidence concerning the timing of Darwin's conversion to the theory of evolution, which, despite recent claims to the contrary, occurred only after his return to England. (shrink)

In 1959, the Charles Darwin Station and the Galápagos National Park were established, formally inaugurating conservation on the archipelago. In the same year, a utopian colony from the United States arrived. Whereas scholars have dismissed the latter and focused on the former, this essay unveils the science-inspired utopianism common to both enterprises. Investing science with the exclusive role of producing all knowledge and steering politics, leaders of the two initiatives aspired not only to protect nature but also to forge (...) a new humanity. Describing how such ambitions burst along lines of race, class, and nationality, I argue that these enterprises consolidated the current understanding of the Galápagos as “pristine”: a site fit for research and tourism but unhospitable to people. Drawing on archival and historical documents, this essay aims to reinvigorate two conversations: one between science and technology studies and conservation, and the other between STS and utopian studies. If recent attempts at bridging the divide between science and imagination have emphasized how powerful actors shape human society, this essay considers the long-lasting effects of scientific imaginaries on a politics of nature. (shrink)

Many environmentalists hold that the loss of a species is intrinsically bad, and many also think that we have moral obligations to species as such. In an attempt to capture these thoughts, some philosophers have suggested that species are bearers of intrinsic value. This approach works well in paradigmatic cases. However, it begins to break down in more difficult scenarios, such as when species boundaries are unclear or when resources are scarce. The case study of the Galápagos giant tortoises (...) in this essay illustrates the limitations of traditional accounts of the non-instrumental value of species. Careful attention to this case indicates how species-centric accounts diverge from the evaluative attitudes of environmentalists, and suggests new directions for theoretical replacements. (shrink)

Though critics have noted the evolutionary themes in Kurt Vonnegut's Galapagos, none has discussed the name Vonnegut gave to the fictional island where most of the novel's action takes place: Santa Rosalia. Since Vonnegut had been reading up on evolutionary ideas while writing the novel, it seems likely that his source for the name was ecologist G. Evelyn Hutchinson's famous 1959 article, “Homage to Santa Rosalia: Or Why Are There So Many Kinds of Animals?” In a study of water bugs (...) in a spring below a cave on Mount Pellegrino where Saint Rosalia's bones had been found in the seventeenth century, Hutchinson established ecological principles regarding speciation, ecological niches, and biodiversity, and he expressed prescient concerns about human impacts on the natural world. These concerns all surface in Galapagos. Of particular interest to Vonnegut must have been Hutchinson's designation of Saint Rosalia as the “patroness of evolutionary studies.”. (shrink)

This study examined knowledge of sea lions for both residents and tourists on San Cristóbal Island in the Galápagos, a famous nature tourism destination. Participants obtained through convenience and snowball sampling answered questionnaires about their knowledge of sea lions. Participants with higher education received higher overall scores, but participants’ education and age influenced answers on only a few questions. Residents and tourists obtained comparable overall scores, exhibiting extensive knowledge of sea lion behavior and life history. Whether participants were residents (...) or tourists influenced answers to several questions, but when only participants with 13 years of education or more were examined, few differences in answers remained between residents or tourists. Participants’ broad knowledge of sea lions may be attributed to the items of knowledge tested, participants’ motivations for travel to the Galápagos, and the fact that sea lions are an engaging and ubiquitous animal. (shrink)

Based on the French notion of terroir or ‘the taste of place,’ a certified geographical indication (GI) identifies an agro-food product as originating in a particular territory and suggests that its quality, reputation, or other characteristics are essentially or exclusively attributable to its geographical origin. Previous scholarship exploring the social construction of terroir has focused on how disparities in political, economic, and cultural power shape GI regulations, certification procedures, and territorial boundaries. While these works have considered knowledge as a resource (...) deployed through relations of authority, studies of GI implementation have not adequately considered an important aspect of power in contemporary politics: the epistemic authority to assert the legitimacy of knowledge and its relevance to policymaking. In contrast, in this article, I take the accomplishment of epistemic authority – to determine the ‘essential’ and ‘exclusive’ physical and/or cultural attributes of place that shape product character – as key to the social construction of terroir and the institutionalization of GI regulations. This process is explored through a case study of Ecuador’s 2013-2019 implementation of a GI for Galápagos Islands coffee. I draw on analysis of relevant policy and regulatory documents and semi-structured interviews with 21 key stakeholders to argue that analytical attention to the legitimacy and relevance of terroir knowledge explains how coffee producers were able to deploy authoritative knowledge to disrupt and reinforce relations of authority and challenge the terms and mechanisms of GI qualification. (shrink)

A Review of Darwin in Galápagos: Footsteps to a New World by K. Thalia Grant and Gregory B. Estes, [2009].

In 1749, Linnaeus presided over the dissertation "Oeconomia Naturae," which argued that each creature plays an important and particular role in nature 's economy. This phrase should be familiar to readers of Darwin, for he claims in the Origin that "all organic beings are striving, it may be said, to seize on each place in the economy of nature." Many scholars have discussed the influence of political economy on Darwin's ideas. In this paper, I take a different tack, showing that (...) Darwin's idea of an economy of nature stemmed from the views of earlier naturalists like Linnaeus and Lyell. I argue, in the first section of the paper, that Linnaeus' idea of oeconomia naturae is derived from the idea of the animal economy, and that his idea of politia naturae is an extension of the idea of a politia civitatis. In the second part, I explore the use of the concept of stations in the work of De Candolle and Lyell – the precursor to Darwin's concept of places. I show in the third part of the paper that the idea of places in an economy of nature is employed by Darwin at many key points in his thinking: his discussion of the Galapagos birds, his reading of Malthus, etc. Finally, in the last section, I demonstrate that the idea of a place in nature 's economy is essential to Darwin's account of divergence. To tell his famous story of divergence and adaptation, Darwin needed the economy of nature. (shrink)

Darwin's theory of evolution brought to an end the static view of nature. It was no longer possible to think of species as immortal, with secure places in nature. Fluctuation of population could no longer be thought of as occurring within definite limits which had been set at the time of creation. Nor was it any longer possible to generalize from the differential reproductive potentials, or from a few cases of mutualism between species, that everything in nature was “fitted to (...) produce general ends, and reciprocal uses.” 134 The appeal to “design” could no longer be substituted for answers to questions concerning animal demography. Instead, the dynamics of a population had to be viewed as the outcome of species' struggle against animate and inanimate factors in the environment. Both the members of a species and the environmental factors tend to vary randomly, and therefore neither evolution nor population dynamics could be fully understood alone. For this reason Darwin's linking of the two subjects was inevitable and not merely an historical accident. Since Darwin had shown that no automatic equilibrium existed, he demonstrated the importance of closer study of the causes of population dynamics and extinction. He also indicated that an understanding of population depends upon the development of a broad knowledge in ecology.Viewed from another direction, Darwin's work ended the early modern era of population studies by clarifying three interrelated problems which were important for understanding population: extinction, distribution, and the nature of species. The components of his answer had been discussed in the eighteenth century, but there had not existed enough evidence for the completion of the revolution in thought which had then begun. At the beginning of the nineteenth century, Playfair found the evidence for extinction conclusive, and, in spite of Lamarck, Curvier convinced the scientific world that there could no longer be any doubt about it. This was a step the importance of which, with his limited knowledge of biogeography and population, Cuvier could not have fully realized. Lamarck attempted, with his evolutionary theory, to circumvent the necessity for admitting extinction, but he overestimated the adaptability of organisms and in doing so he underestimated the importance of competition and the whole field of ecology. On the other hand, he was not willing to let questions such as the origin of species remain taboo to science. The origin of species was a biogeographical as well as a paleontological question. Humboldt correlated environment with the distribution of species and conveyed the impression that plant communities are subject to change. De Candolle, following the lead of Linnaeus and Humboldt, emphasized the ecological aspects of biogeography, not only the importance of habitat and range, clearly showing the ecological effects of competition. The entomologists Kirby and Spence took a faltering step toward understanding the relationship between population and ecological role, but they fell short of any significant new conclusions. Neither they nor Swainson could fully comprehend the new perspective of De Candolle.Lyell was able to bring together the evidence from these three lines of investigation and weave them into an important synthesis that almost accomplished that Darwin later did. Although opposing Lamarck's theory of evolution, Lyell had a dynamic view of ecology. He realized that population dynamics offered an important key to the understanding of biogeography. Since he knew that species become extinct, he investigated closely the factors which could either preserve or extinguish species. While explaining these factors, he described the interrelationships of species in greater detail than had ever been done before. Forbes continued to develop Lyell's ecological concepts, and his first-hand field experience enabled him to describe biotic communities more concretely than Lyell had.Having the advantages of Lyell's understanding and his own experience from a global voyage, Darwin could take the final step from the static to the dynamic concept of life. He had seen populations fluctuating and also fossil species in South America, and on the Galapagos Islands he had encountered a biogeographical problem that could not be credibly solved without the idea of evolution. However, the bare idea of evolution did not fully answer his questions. He sought physiological causes of extinction before he read Malthus and realized that De Candolle and Lyell had correctly emphasized the importance of competition. Darwin found that, in order to understand evolution, he needed to improve his understanding of ecology. He wanted to know when populations were most easily decimated, how extensive were competition and cooperation, what effects parasites have upon populations, and what changes occur in biotic communities when a species is either added or subtracted. He contributed to some extent to answering these questions. Though there remained much for others to do, there was now a new and more secure theoretical framework within which later studies could be interpreted. As Ernst Mayr has observed, Darwin's “consistent thinking in terms of population has had an impact on biological theory and practice which is second only to his sponsorship of natural selection as the mechanism of evolution.” 135. (shrink)

Overview * Part I: Introduction * Philip Appleman, Darwin: On Changing the Mind * Part II: Darwin’s Life * Ernst Mayr, Who Is Darwin? * Part III: Scientific Thought: Just before Darwin * Sir Gavin de Beer, Biology before the Beagle * Thomas Robert Malthus, An Essay on the Principle of Population * William Paley, Natural Theology * Jean Baptiste Pierre Antoine de Monet Lamarck, Zoological Philisophy * Charles Lyell, Principles of Geology * John Herschell, The Study of Natural Philosophy (...) * William Whewell, Astronomy and General Physics Considered with Reference to Natural Theology * Alfred Russel Wallace, On the Tendency of Varieties to Depart Indefinitely from the Original Type * Part IV: Selections from Darwin’s Work * The Voyage of the Beagle * o Chapter I. St. Jago-Cape de Verd Island o Chapter XVII. Galapagos Archipelago * On the Tendency of Species to Form Varieties; and On the Perpetuation of Varieties and Species by Natural Means of Selection * o I. Extract from an unpublished Work on Species, by C. Darwin, Esq.... o II.of Letter from C. Darwin, Esq., to Prof. Asa Gray, Boston, U.S., dated Down, September 5th, 1857 * An Historical Sketch of the Progress of Opinion on the Origin of Species, previously to the : Publication of This Work The Origin of Species * o Introduction o Chapter I. Variation under Domestication o Chapter II. Variation under Nature o Chapter III. Struggle for Existence o Chapter IV. Natural Selection o Chapter VI. Difficulties on Theory o Chapter IX. On the Imperfections of the Geological Record o Chapter XIII. Mutual Affinities of Organic Beings: Morphology: Embryology: Rudimentary Organs o Chapter XIV. Recapitulation and Conclusion * The Descent of Man * o Introduction o Chapter I. The Evidence of the Descent of Man from Some Lower Form o Chapter II. On the Manner of Development of Man from Some Lower Form o Chapter III. Comparison of the Mental Powers of Man and the Lower Animals o Chapter VI. On the Affinities and Genealogy of Man o Chapter VIII. Principles of Sexual Selection o Chapter XIX. Secondary Sexual Characters of Man o Chapter XX. Secondary Sexual Characters of Man-continued o Chapter XXI. General Summary and Conclusion * Part V: Darwin’s Influence on Science * THE VICTORIAN OPPOSITION TO DARWIN * o David L. Hull, Darwin and His Critics o Adam Sedgwick, Objections to Mr. Darwin’s Theory of the Origin of Species o Sir Richard Owen, Darwin on the Origin of Species o Fleeming Jenkin, Review of the Origin of Species * VICTORIAN SUPPORTERS OF DARWIN * o Joseph Dalton Hooker, Flora Tasmaniae o Thomas Henry Huxley, On the Relations of Man to the Lowe Animals o Charles Lyell, Principles of Geology o Alfred Russel Wallace, The Debt of Science to Darwin * DARWIN AND THE SHAPING OF MODERN SCIENCE * o Scientific Method in Evolution o National Academy of Sciences, Evolution and the Nature of Science o Richard Dawkins, Explaining the Very Improbable o Lewis Thomas, On the Uncertainty of Science o Noretta Koetge, Postmodernisms and the Problem of Scientific Literary o Richard Dawkins, Science and Sensibility o The Neo-Darwinian Synthesis o Peter Bowler, The Evolutionary Synthesis o The Human Genealogy o Adam Kuper, The Chosen Primate o Ian Tattersall, Out of Africa Again... and Again? o Stephen Jay Gould, The Human Difference o Punctuated Equilibrium o Stephen Jay Gould, [On Punctuated Equilibrium] o Niles Eldredge, The Great Stasis Debate o Rethinking Taxonomy o Kevin Padian, Darwin’s Views of Classification o David L. Hull, Cladistic Analysis o Kevin Padian and Luis M. Chiappe, Cladistics in Action: The Origin of Birds and Their Flight o Evolution as Observable Fact o James L. Gould and William T. Keeton with Carol Grant Gould, How Natural Selection Operates o Peter r. Grant, Natural Selection and Darwin’s Finches o John A. Endler, Natural Selection in the Wild * Part VI: Darwinian Patterns in Social Thought * COMPETITION AND COOPERATION * o Richard Hofstadter, The Vogue of Spencer o Andrew Carnegie, The Gospel of Wealth o Peter Kropotkin, Mutual Aid o Martin A. Nowak, Robert M. May, and Karl Sigmund, The Arithmetics of Mutual Help * NATURE AND NURTURE * o Edward O. Wilson, Sociobiology: The New Synthesis o Stephen Jay Gould, Biological Potentiality vs. Biological Determination o Barbara Ehrenreich and Janet McIntosh, The New Creationism: Biology under Attack * EVOLUTION AND GENDER * o Elizabeth Cady Stanton, The Woman’s Bible o Nancy Makepeace Tanner, On Becoming Human o Evelleen Richards, Darwin and the Descent of Woman o James Eli Adams, Woman Red in Tooth and Claw * EVOLUTION AND OTHER DISCIPLINES * o Edward O. Wilson, [On Consilience] o Randolph H. Nesse and George C. Williams, Evolution and the Origin of Disease o Steven Pinker, How the Mind Works o Steve Jones, The Set within the Skull * Part VII: Darwinian Influences in Philosophy and Ethics * John Dewey, The Influence of Darwin on Philosophy * Daniel C. Dennett, Darwin’s Dangerous Idea: Natural Selection as an Algorithmic Process * Michael Ruse Darwinian Epistemology * Thomas Henry Huxley, Evolution and Ethics * Julian Huxley, Evolutionary Ethics * Michael Ruse and Edward O. Wilson, The Evolution of Ethics * Frans de Waal, Good Natured: The Origin of Right and Wrong in Humans and Other Animals * Matt Ridley, The Origins of Virtue * Part VIII: Evolutionary Theory and Religious Theory * MAINSTREAM RELIGIOUS SUPPORT FOR EVOLUTION * o Pope John Paul II, Message to the Pontifical Academy of Sciences o Central Conference of American Rabbis, On Creationism in School Textbooks o United Presbyterian Church in the U.S.A., Evolution and Creationsim o The Lutheran World Federation, [Statement on Evolution] o The General Convention of the Episcopal Church, Resolution on Evolutionism and Creationism o Unitariuan Universalist Association, Resolution Opposing "Scientific Creationism" * FUNDAMENTALIST CREATIONISM * o Eugene C. Scott, Antievolution and Creationism in the United States o The Scopes Trial o Thomas McIver, Orthodox Jewish Creationists o Harun Yahya, [Islamic Creationism] o Seami Srila Prabhupada, [A Hare Krishna on Darwinian Evolution] o Institute for Creation Research, Tenets of Creationism o Henry M. Morris, Scientific Creationism o Thomas J. Wheeler, Review of Morris o Richard D. Sjolund and Betty McCollister, Evolution at the Grass Roots o Richard D. Sjolund, [Creationism versus Biotechnology] o Betty McCollister, [The Politics of Creationism] o Molleen Matsumara, What Do Christians Really Believe about Evolution? o National Center for Science Education, Seven Significant Court Decisions Regarding Evolution/Creation Issues * PERSONAL INCREDULITY AND ANTIEVOLUTIONISM * o Richard Dawkins, [The Argument from Personal Incredulity] o Phillip E. Johnson, Darwin on Trial o Eugenie C. Scott, Review of Johnson o Michael Behe, Darwin’s Black Box o Robert Dorit, Review of Behe o Michael Ruse, Darwin’s New Critics on Trial * SCIENTISTS’ OPPOSITION TO CREATIONISM * o American Association for the Advancement of Science, Forced Teaching of Creationist Beliefs in Public School Science Education o American Institute of Biological Sciences, Resolution Oposing Creationism in Science Courses o National Association of Biology Teachers, Statement on Teaching Evolution o National Academy of Sciences, Frequently Asked Questions about Evolution and the Nature of Science * FUNDAMENTALIST CREATIONISM AND THE VALUE OF SATIRE * o Michael Shermer, Genesis Revisted: A Scientific Creation Story o Philip Appleman, Darwin’s Ark * Part IX: Darwin and the Literary Mind * DARWIN’S LITERARY SENSIBILITY * o Charles Darwin, Autobiography o L. Robert Stevens, Darwin’s Humane Reading o George Levine, Darwin and Pain: Why Science Made Shakespeare Nauseating o Gillian Beer, Darwin’s Plots * DARWIN’S INFLUENCE ON LITERATURE * o Lionel Stevenson, Darwin among the Poets o George Levine, Darwin among the Novelists o Joseph Wood Krutch, The Tragic Fallacy o Herbert J. Muller, Modern Tragedy o Philip Appleman, Darwin-Sightings in Recent Literature. (shrink)

Adaptation by means of natural selection depends on the ability of populations to maintain variation in heritable traits. According to the Modern Synthesis this variation is sustained by mutations and genetic drift. Epigenetics, evodevo, niche construction and cultural factors have more recently been shown to contribute to heritable variation, however, leading an increasing number of biologists to call for an extended view of speciation and evolution. An additional common feature across the animal kingdom is learning, defined as the ability to (...) change behavior according to novel experiences or skills. Learning constitutes an additional source for phenotypic variation, and change in behavior may induce long lasting shifts in fitness, and hence favor evolutionary novelties. Based on published studies, I demonstrate how learning about food, mate choice and habitats has contributed substantially to speciation in the canonical story of Darwin’s finches on the Galapagos Islands. Learning cannot be reduced to genetics, because it demands decisions, which requires a subject. Evolutionary novelties may hence emerge both from shifts in allelic frequencies and from shifts in learned, subject driven behavior. The existence of two principally different sources of variation also prevents the Modern Synthesis from self-referring explanations. (shrink)

“I often said before starting, that I had no doubt I should frequently repent of the whole undertaking.” So wrote Charles Darwin aboard The Beagle , bound for the Galapagos Islands and what would arguably become the greatest and most controversial discovery in scientific history. But the theory of evolution did not spring full-blown from the head of Darwin. Since the dawn of humanity, priests, philosophers, and scientists have debated the origin and development of life on earth, and with modern (...) science, that debate shifted into high gear. In this lively, deeply erudite work, Pulitzer Prize–winning science historian Edward J. Larson takes us on a guided tour of Darwin’s “dangerous idea,” from its theoretical antecedents in the early nineteenth century to the brilliant breakthroughs of Darwin and Wallace, to Watson and Crick’s stunning discovery of the DNA double helix, and to the triumphant neo-Darwinian synthesis and rising sociobiology today. Along the way, Larson expertly places the scientific upheaval of evolution in cultural perspective: the social and philosophical earthquake that was the French Revolution; the development, in England, of a laissez-faire capitalism in tune with a Darwinian ethos of “survival of the fittest”; the emergence of Social Darwinism and the dark science of eugenics against a backdrop of industrial revolution; the American Christian backlash against evolutionism that culminated in the famous Scopes trial; and on to today’s world, where religious fundamentalists litigate for the right to teach “creation science” alongside evolution in U.S. public schools, even as the theory itself continues to evolve in new and surprising directions. Throughout, Larson trains his spotlight on the lives and careers of the scientists, explorers, and eccentrics whose collaborations and competitions have driven the theory of evolution forward. Here are portraits of Cuvier, Lamarck, Darwin, Wallace, Haeckel, Galton, Huxley, Mendel, Morgan, Fisher, Dobzhansky, Watson and Crick, W. D. Hamilton, E. O. Wilson, and many others. Celebrated as one of mankind’s crowning scientific achievements and reviled as a threat to our deepest values, the theory of evolution has utterly transformed our view of life, religion, origins, and the theory itself, and remains controversial, especially in the United States (where 90% of adults do not subscribe to the full Darwinian vision). Replete with fresh material and new insights, Evolution will educate and inform while taking readers on a fascinating journey of discovery. (shrink)

Gathers selections from nine of Darwin's most important books, including writings about coral reefs, the Galapagos Islands, evolution, emotions, and flowers.

This article takes as its starting point and its main problematic the status of evolution as a ‘sterile belief’ in contemporary technoscientific culture. Focusing in particular on the role of evolution across the boundaries of art and science in the contexts of artificial life and transgenic engineering, it offers a critique of the belief in evolutionary possibility as an abstract process. The lack of what François Jacob refers to as a dialogue between the possible and the actual is seen to (...) account for the sterility of evolutionary possibility and for its status as myth. In particular, evolutionary possibility recalls the myth of metamorphoses, the conservatism of which is demonstrated in an analysis of two art/science works - Galápagos by Karl Sims and Genesis by Eduardo Kac. Lacking a dialogue with the actual, metamorphic evolutionary possibility here becomes self-referential and produces nothing new. Rather than a rejection of evolution per se, this article takes up Jacob’s critique of its over-extension by showing how this operates in two works representing the fields of artificial life and transgenesis respectively. (shrink)

The standard picture of evolution, is externalist: a causal arrow runs from environment to organism, and that arrow explains why organisms are as they are (Godfrey-Smith 1996). Natural selection allows a lineage to accommodate itself to the specifics of its environment. As the interior of Australia became hotter and drier, phenotypes changed in many lineages of plants and animals, so that those organisms came to suit the new conditions under which they lived. Odling-Smee, Laland and Feldman, building on the work (...) of Richard Lewontin, have shown that while sometimes appropriate, this is an inadequate conception of the relationship between organisms and the environments in which they live. Over time organisms alter their environment as well as being altered by their environments (Lewontin 1982; Lewontin 1983; Lewontin 1985). For example, animals modulate the effects of their physical and biological environment by building shelters: the beaver’s dam and lodge system, and termite mounds are two famous cases of animal structures, but they are few of many. There are many thousands of animals which make nests, burrows and other shelters. Likewise, animals make tools that give them access to resources from which they would otherwise be excluded: thus the Galapagos woodpecker finch uses a cactus needle to extract insects from crevasses in bark — insects that they would otherwise be unable to catch (Tebbich, Taborsky et al. 2001). Tool making is not as common as shelter-making, but it is common. For example many animals make traps: there are many species of pit-making antlions. Thus in part organisms make the world in which they live. They partially construct their own niches. Odling-Smee, Laland and Feldman argue that this has five major and under-appreciated consequences for biological theory. (shrink)

The value of extending the human lifespan remains a key philosophical debate in bioethics. In building a case against the extension of the species-typical human life, Nicolas Agar considers the prospect of transforming human beings near the end of their lives into Galapagos tortoises, which would then live on decades longer. A central question at stake in this transformation is the persistence of human consciousness as a condition of the value of the transformation. Agar entertains the idea that consciousness could (...) persist in some measure, but he thinks little is to be gained from the transformation because the experiences available to tortoises pale in comparison to those available to human beings. Moreover, he thinks persisting human consciousness and values would degrade over time, being remade by tortoise needs and environment. The value available in the transformation would not, then, make the additional years of life desirable. Agar’s account does not, however, dispose of the tortoise transformation as a defensible preference. Some people might still want this kind of transformation for symbolic reasons, but it would probably be better that no continuing human consciousness persist, since that consciousness would be inexpressible as such. Even so, it is not irrational to prefer various kinds of lifespan extension even if they involve significant modifications to human consciousness and values. (shrink)

The value of extending the human lifespan remains a key philosophical debate in bioethics. In building a case against the extension of the species-typical human life, Nicolas Agar considers the prospect of transforming human beings near the end of their lives into Galapagos tortoises, which would then live on decades longer. A central question at stake in this transformation is the persistence of human consciousness as a condition of the value of the transformation. Agar entertains the idea that consciousness could (...) persist in some measure, but he thinks little is to be gained from the transformation because the experiences available to tortoises pale in comparison to those available to human beings. Moreover, he thinks persisting human consciousness and values would degrade over time, being remade by tortoise needs and environment. The value available in the transformation would not, then, make the additional years of life desirable. Agar’s account does not, however, dispose of the tortoise transformation as a defensible preference. Some people might still want this kind of transformation for symbolic reasons, but it would probably be better that no human consciousness persist, since that consciousness would be inexpressible as such. Even so, it is not irrational to prefer various kinds of lifespan extension even if they involve significant modifications to human consciousness and values. (shrink)

A recent analysis of the genomes of Darwin's finches revealed extensive interspecies allele sharing throughout the history of the radiation and identified a key locus responsible for morphological evolution in this group. The radiation of Darwin's finches on the Galápagos archipelago has long been regarded as an iconic study system for field ecology and evolutionary biology. Coupled with an extensive history of field work, these latest findings affirm the increasing acceptance of introgressive hybridization, or gene flow between species, as (...) a significant contributor to adaptive evolution. Here, we review and discuss these findings in relation to both classical work on Darwin's finches and contemporary work showing similar evolutionary signatures in other biological systems. The continued unification of genomic data with field biology promises to further elucidate the molecular basis of adaptation in Darwin's finches and well beyond. (shrink)

In lieu of an abstract, here is a brief excerpt of the content:Five Poems AMIT MAJMUDAR Observing Orpheus I hear the meaning turn back in his throat like Eurydice on the way up from the darkness. Music’s meaning is its making. As for me, I am one more animal in his entourage, learning a new thirst, finding a new south. None of us knew we had this instinct in us. If deserts hide wildflowers until first rain, bright ears are blossoming (...) out of our skulls. He doesn’t have much longer. I know this myth. A God douses the fire with a beehive. The Maenads smear their faces with a warpaint made of stoneground fireflies and pine sap. Hands—hands like his, that drew music through lyre strings into our forest like pieces of bread through a prison fence— are reaching for his body now, his lyre. Their weapons are hands, nothing but hands. They are infinite. They are enough. arion 27.1 spring/summer 2019 Her Metamorphoses First a beetle, body armor forged of chitin, exoskeleton he’ll never bruise, Undiscovered, unapologetically hermetic on her own Galapagos. Next a fruit bat, echolocating tangerines like a blind woman singing up at stars. Then a patio lizard, sunning herself for as long as she needs, A better place for keeping warm than bed with him. Next a sleek water moccasin, quick to make out another woman’s heat signature on his neck, Even quicker to strike. A caterpillar to poke holes in his lying roses And from that destruction to fashion two beautiful wings for herself. A buzzard, to clean up the roadside pile he left of her. A woman in full with no more soft blind pupa stages of her life cycle left, No more forms to force herself to fit for the sake of his pleasure, No more shapeshifts, no more slimming stripes, no more abdominal crunches: Just this body, just this last name, just this apartment, just this bowl of midnight cereal. Just her. 106 five poems Sisyphus Shove hard, shove, shove hard, eyes bloodshot, knees bent Till your boulder goes weightless and races ahead in descent. Force its dark door open inch by grim inch, Shove hard, shove, shove hard, eyes bloodshot, knees bent, Heels dug deep in hell’s steep hillside, teeth clenched. What gravity strips from you, gravity grants you again. Shove hard, shove, shove hard, eyes bloodshot, knees bent Till your body goes weightless and races ahead in ascent. Amit Majmudar 107 A Trinket for Persephone A deathwatch beetle burrows in the pale hall of your marrow, your underground, your tube of bone on which you’re riding home alone. This blackwinged locket with your soul emerges rattling from a hole, a rattle in your emptied throat, the diva’s final, graceless notes, a clicking echoed in the rafters, this bugbear insect coming after to scuttle across the marshy chest, sinkhole navel, and pubic nest out to the yard, below the porch untrackable by any torch until this bug with you inside has burrowed to the other side. It drops into the underworld, as if from some cocoon unfurled, to scale a queenly wrist and rest your brooch of black upon her breast. 108 five poems Four Ways of Looking at Argus 1. His Vigil These hundred eyes, these hundred balloons Tied to the boy-small wrist of my mind Are down to ninety-nine. I lost one to a bee sting late last June— I never thought of them as fragile Until that pinprick broke my vigil. I’m slowing down. I didn’t blink in time. I have no inkling why I keep This vigil—surely not to watch, Panoptic from this mountaintop, A heifer drifting off to sleep? I’m here, I must be here, to see A signal fire from the east That warns of Olympus under siege And Ares gory from a rout, Athena’s owl’s eyes gouged out, The naiads naked in a pen, The whole world given up to men. The Gods will need a monster then. 2. After Martial Hermes was in a fix: He needed lullabies To shutter Argus’s one hundred eyes And get the tethered heifer—coal-eyed Io—free Of Hera’s... (shrink)

The aim of this paper is to show how James Mark Baldwin’s theory of Organic Selection can be fruitfully integrated with Alfred North Whitehead’s speculative philosophy, as part of the endeavor to develop a comprehensive process-relational evolutionary cosmology. In so doing, it provides an overview of the theory of Organic Selection and points to several concrete examples from the Galapagos Islands which elucidate Baldwin’s claim that organisms, through their selective activities and behavioral adjustments, play a causal role in directing evolutionary (...) processes. I emphasize some of the affinities between Baldwin’s theory of Organic Selection and Whitehead’s theory of prehensions, especially focusing on the latter’s notion of “prehensive selectivity.” Overall, while Baldwin’s theory of Organic Selection provides a biological ground for a comprehensive process-relational evolutionary cosmology to be developed, illuminating the importance of Whitehead’s theory of prehensions for evolutionary theory, Whitehead’s overall speculative scheme can, in turn, strengthen the metaphysical, epistemological, and ethical foundations of Baldwin’s theory. In the course of merging the two views, I arrive at an enlarged conception of Organic Selection, placing it in context with Darwin’s principle of Natural Selection. At the end of the paper, I take up the resulting question of the ethics of selectivity in general, arguing that the merger of Baldwin’s and Whitehead’s ideas constitutes a “non-reductionistic critical pan-selectionism.” This position stands in contrast to the antagonistic standpoints of “Selectionism” and “Anti-Selectionism” in the ongoing debates over the ethical dimensions of Darwinian evolution. (shrink)

A flurry of technological advances in molecular, cellular and developmental biology during the past decade has provided a clearer understanding of mechanisms underlying phenotypic diversification. Building upon such momentum, a recent paper tackles one of the foremost topics in evolution, that is the origin of species‐specific beak morphology in Darwin's finches.1 Previous work involving both domesticated and wild birds implicated a well‐known signaling pathway (i.e. bone morphogenetic proteins) and one population of progenitor cells in particular (i.e. cranial neural crest), as (...) primary factors for establishing beak size and shape. But these results were limited in their ability to explain fully the morphogenetic bases of patterned outgrowth. So in a quest to identify novel genes whose expression correlated with differences in beak anatomy among Darwin's finches, a DNA microarray approach was undertaken using tissues harvested from the Galápagos Islands. The results are striking and point to a protein called calmodulin, which is a mediator of cellular calcium signaling, as a key determinant of beak length. BioEssays 29: 1–6, 2007. © 2006 Wiley Periodicals, Inc. (shrink)

What role can technology play in cultivating a disciplinary stance — raising questions, planning investigations, interpreting data and constructing explanations in a way that reflects disciplinary values and principles? How can overt and tacit expert scientific knowledge be captured, represented and used to design software that enables novices to assume a disciplinary stance in their investigations? We present The Galapagos Finches software designed to foster a biological and evolutionary stance. Our approach, Discipline-Specific Strategic Support , translates the main variable types, (...) comparison types and relationships in a discipline into manipulable objects in the interface. Pre/post-tests show how DSSS helps achieve a balance between content and process goals. A contrastive-case microanalysis of high, medium and low-achieving students' inquiry shows progress toward a disciplinary stance. Our study shows how software representations carry multiple levels of meaning, and that the efficacy of learning technologies hinges on reflection at both the navigation and disciplinary-signification levels. (shrink)

Discoverer of the theory of evolution by natural selection, and thus the founder of modern evolutionary biology, Charles Darwin is responsible for one of the most fundamental and far‐reaching contributions to the modern scientific world view. Born in 1809 in Shrewsbury into a wealthy Victorian family, Darwin was educated at the universities of Edinburgh and Cambridge. Though his formal education was of little interest to him ‐ “my time was wasted, as far as the academical studies were concerned” (1969, p. (...) 58) ‐ Cambridge did provide the opportunity to pursue his childhood interest in natural history. Upon graduating in 1831, Darwin joined a charting expedition around South America on board HMS Beagle, which was to last for five years. During this time, he made lengthy expeditions inland, studying the flora, fauna, and geology of South America and the Pacific Islands, and accumulating a massive biological and geological collection. These years were critical for Darwin's intellectual development, as he admitted, sowing the seeds of his doubt in the fixity of individual species. Darwin was impressed by a large number of facts that could not be explained on the assumption of the fixity of species, including the close structural similarities between extinct and living species revealed by the fossil record. Most important of all were the tortoises and finches of the Galapagos Islands: each island was inhabited by very similar, yet distinct varieties ‐ clear evidence for Darwin of common ancestry shaped by adaptation to local conditions. The creatures of the Galapagos, Darwin wrote, bring us close “to that great fact ‐ that mystery of mysteries ‐ the first appearance of new beings on this earth” (1839, p. 466). (shrink)

What role can technology play in cultivating a disciplinary stance — raising questions, planning investigations, interpreting data and constructing explanations in a way that reflects disciplinary values and principles? How can overt and tacit expert scientific knowledge be captured, represented and used to design software that enables novices to assume a disciplinary stance in their investigations? We present The Galapagos Finches software designed to foster a biological and evolutionary stance. Our approach, Discipline-Specific Strategic Support, translates the main variable types, comparison (...) types and relationships in a discipline into manipulable objects in the interface. Pre/post-tests show how DSSS helps achieve a balance between content and process goals. A contrastive-case microanalysis of high, medium and low-achieving students' inquiry shows progress toward a disciplinary stance. Our study shows how software representations carry multiple levels of meaning, and that the efficacy of learning technologies hinges on reflection at both the navigation and disciplinary-signification levels. (shrink)