The work of Pere Alberch is crucial to study the early stages of evo-devo. In particular, it illustrates very persuasively why developmental systems have so much to say about the course of evolutionary change. In addition to an important empirical work, he elaborated a stimulating framework of theoretical ideas on biological form, morphological variation, and how developmental processes establish possible evolutionary paths previous to the action of natural selection. In this framework, the study of development and evolution are related through (...) the notion of possible morphologies. In his view, the morphology of organisms shows internal coherence and structure, emergent from complex non linear interactions among parts and with the environment. (shrink)
In the Modern Synthesis the study of patterns refers to how to identify and systematize order in lineages (description), attributed to underlying processes or mechanisms (explanation). But patterns and processes play distinct roles in evodevo. In this paper we (1) distinguish three different views (the transformational, the morphogenetic and the process approach) according to the role they play in the description and explanation of development and evolution, and (2) relate this discussion to the issues of homology and variation.
In the evolutionary biology of the Modern Synthesis the study of patterns refers to how to identify and systematise order in lineages, looking for hierarchies or for branching/splitting events in the tree of life, whereas the resulting order is supposed to be due to underlying processes or mechanisms. But patterns and processes play distinct roles in evo-devo: four different views on the role of patterns and processes in descriptions and explanations of development and evolution: A) transformational; B) generative; C) processual; (...) and D) complex are reviewed in this paper. Then, this discussion is related to two issues in evo-devo: homology and variation. (shrink)
The notion of fitness is usually equated to reproductive success. However, this actualist approach presents some difficulties, mainly the explanatory circularity problem, which have lead philosophers of biology to offer alternative definitions in which fitness and reproductive success are distinguished. In this paper, we argue that none of these alternatives is satisfactory and, inspired by Mumford and Anjum’s dispositional theory of causation, we offer a definition of fitness as a causal dispositional property. We argue that, under this framework, the distinctiveness (...) that biologists usually attribute to fitness—namely, the fact that fitness is something different from both the physical traits of an organism and the number of offspring it leaves—can be explained, and the main problems associated with the concept of fitness can be solved. Firstly, we introduce Mumford and Anjum's dispositional theory of causation and present our definition of fitness as a causal disposition. We explain in detail each of the elements involved in our definition, namely: the relationship between fitness and the functional dispositions that compose it, the emergent character of fitness, and the context-sensitivity of fitness. Finally, we explain how fitness and realized fitness, as well as expected and realized fitness are distinguished in our approach to fitness as a causal disposition. (shrink)
In identifying intrinsic molecular chance and extrinsic adaptive pressures as the only causally relevant factors in the process of evolution, the theoretical perspective of the Modern Synthesis had a major impact on the perceived tenability of an ontology of dispositional properties. However, since the late 1970s, an increasing number of evolutionary biologists have challenged the descriptive and explanatory adequacy of this “chance alone, extrinsic only” understanding of evolutionary change. Because morphological studies of homology, convergence, and teratology have revealed a space (...) of possible forms and phylogenetic trajectories that is considerably more restricted than expected, evo-devo has focused on the causal contribution of intrinsic developmental processes to the course of evolution. Evo-devo’s investigation into the developmental structure of the modality of morphology – including both the possibility and impossibility of organismal form – has led to the utilisation of a number of dispositional concepts that emphasise the tendency of the evolutionary process to change along certain routes. In this sense, and in contrast to the perspective of the Modern Synthesis, evo-devo can be described as a “science of dispositions.” This chapter discusses the recent philosophical literature on dispositional properties in evo-devo, exploring debates about both the metaphysical and epistemological aspects of the central dispositional concepts utilised in contemporary evo-devo and addressing the epistemological question of how dispositional properties challenge existing explanatory models in evolutionary biology. (shrink)
La biología sintética mantiene una relación muy singular con la teoría evolutiva: por un lado, parte de una interpretación ingenieril de la evolución para fundar su aproximación al diseño de bioartefactos; por otro, la biología sintética aspira, en última instancia, a deshacerse de la evolución creando organismos de novo que se comporten de un modo predecible. Tras examinar las tres grandes propiedades que aparecen recurrentemente en la descripción sintética de los nuevos artefactos orgánicos, argumentaré que la biología sintética se erige (...) sobre una concepción de los organismos y en particular de la relación de estos con su legado histórico que ha sido profundamente cuestionada por la biología evolucionista contemporánea. (shrink)
Despite the radical importance of embryology in the development of organicism, developmental biology remains philosophically underexplored as a theoretical and empirical resource to clarify the nature of organisms. This paper discusses how embryology can help develop the organisational definition of the organism as a differentiated, functionally integrated, and autonomous system. I distinguish two conceptions of development in the organisational tradition that yield two different conceptions of the organism: the life-history view claims that organisms can be considered as such during their (...) whole ontogeny; the constitutive view distinguishes two periods in the life history, a period of generation and a period of self-maintenance of a constitutive organisation. Arguing in favour of the constitutive view, it will be claimed that the organisational criteria for the definition of organism (i.e., differentiation, functional integration, and autonomy) can only be applied to the developmental system when it has entered the period of self-maintenance of a constitutive organisation. Under the light of current research in developmental biology, it is possible to make explicit how organisms come to be as organisms. To this end, I explore key ontogenetic events that help us clarify the core aspects of animal organisation and allow us to identify the developmental stage that marks the ontological transition between an organism in potency and an organism in actuality. The structure of this ontogenetic unfolding parallels the conceptual structure of the very notion of organism; the generation of the being of a particular organism parallels its definition. (shrink)
While the notion of chance has been central in discussions over the probabilistic nature of natural selection and genetic drift, its role in the production of variants on which populational sampling takes place has received much less philosophical attention. This article discusses the concept of chance in evolution in the light of contemporary work in evo-devo. We distinguish different levels at which randomness and chance can be defined in this context, and argue that recent research on variability and evolvability demands (...) a causal understanding of variational probabilities under which development acquires a creative, rather than a constraining role in evolution. We then provide a propensity interpretation of variational probabilities that solves a conceptual confusion between causal properties, variational probabilities and extant variation present in the literature, and explore some metaphysical consequences that follow from our interpretation, specifically with regards to the nature of developmental types. 2.1Mutational randomness 2.2Morphological randomness 2.3Adaptive randomness3Variability and Phenotypic Probabilities 3.1The RNA model as a model for evo-devo4Evolvability and the Probability of Evolving5Variational Propensities in Evo-Devo 5.1Individuating variational propensities 5.2Developmental types6Conclusion. (shrink)
Despite the radical importance of embryology in the development of organi- cism, developmental biology remains philosophically underexplored as a theoretical and empirical resource to clarify the nature of organisms. This paper discusses how embryology can help develop the organisational definition of the organism as a differentiated, function- ally integrated, and autonomous system. I distinguish two conceptions of development in the organisational tradition that yield two different conceptions of the organism: the life- history view claims that organisms can be considered as (...) such during their whole ontogeny; the constitutive view distinguishes two periods in the life history, a period of generation and a period of self-maintenance of a constitutive organisation. Arguing in favour of the con- stitutive view, it will be claimed that the organisational criteria for the definition of organ- ism (i.e., differentiation, functional integration, and autonomy) can only be applied to the developmental system when it has entered the period of self-maintenance of a constitutive organisation. under the light of current research in developmental biology, it is possible to make explicit how organisms come to be as organisms. To this end, I explore key onto- genetic events that help us clarify the core aspects of animal organisation and allow us to identify the developmental stage that marks the ontological transition between an organism in potency and an organism in actuality. The structure of this ontogenetic unfolding paral- lels the conceptual structure of the very notion of organism; the generation of the being of a particular organism parallels its definition. (shrink)
Criticisms of the “container” model of pregnancy picturing female and embryo as separate entities multiply in various philosophical and scientific contexts during the last decades. In this paper, we examine how this model underlies received views of pregnancy in evolutionary biology, in the characterization of the transition from oviparity to viviparity in mammals and in the selectionist explanations of pregnancy as an evolutionary strategy. In contrast, recent evo-devo studies on eutherian reproduction, including the role of inflammation and new maternal cell (...) types, gather evidence in favor of considering pregnancy as an evolved relational novelty. Our thesis is that from this perspective we can identify the emergence of a new historical individual in evolution. In evo-devo, historical units are conceptualized as evolved entities which fulfill two main criteria, their continuous persistence and their non-exchangeability. As pregnancy can be individuated in this way, we contend that pregnant females are historical individuals. We argue that historical individuality differs from, and coexists with, other views of biological individuality as applied to pregnancy, but brings forward an important new insight which might help dissolve misguided conceptions. (shrink)
In September 2008, 10 years after the untimely death of Pere Alberch (1954–1998), the 20th Altenberg Workshop in Theoretical Biology gathered a group of Pere’s students, col- laborators, and colleagues (Figure 1) to celebrate his contribu- tions to the origins of EvoDevo. Hosted by the Konrad Lorenz Institute for Evolution and Cognition Research (KLI) outside Vienna, the group met for two days of discussion. The meeting was organized in tandem with a congress held in May 2008 at the Cavanilles Institute (...) for Biodiversity and Evolutionary Biology (ICBiBE) in Valencia, Spain. The talks at the KLI were equal parts: nostalgic remembrance, excitement over new ways of thinking about old problems, and an unrepressed vitriol against the resurgence of reductionist thinking in EvoDevo. Here we highlight some of the key aspects of Pere’s life and work that informed and infused the talks. (shrink)
The views of Celia Amorós, which are dealt with here, originate in the ideas which appeared, especially in the Enlightenment, and which demanded equality for women. Following this line of thought she studies some of the most powerful theories of Western philosophical tradition in order to carry out a critical feminist deconstruction. Since her work is a critique of the philosophical critique of the Enlightenment, one can identify traces , today almost forgotten, of a feminist tradition of thought which can (...) be considered as a discourse of emancipation. Furthermore, this critical exercise becomes an intellectual and social non-patriarchal project of a society free from gender restrictions. In this sense, Celia Amoros’ philosophical work develops into philosophical feminism, not only in terms of being critical revision of the history of ideas, but also and principally because it takes shape as critical reflection and as direct dialogue with present-day philosophical discourse. (shrink)
En los años �30 del siglo pasado surge La Legión de la Decencia, en Estados Unidos, radicalizando una ola inquisitorial ya iniciada años atrás para imponer la censura en el cine. Este artículo se sumerge en ese Hollywood en vigilancia permanente y fija su atención en las mujeres de aquella industria: directoras, guionistas, editoras, actrices� Profesionales de valía excepcional que sufrieron los rigores de los centinelas de la moral. Un plano panorámico, con un enfoque feminista, que nos recuerda algunos nombres, (...) títulos y fechas inolvidables de ese universo cinematográfico. (shrink)
Este artículo estudia los elementos de estrategia comercial y comunicativa queintervienen en las interacciones producidas en espacios de consumo. Este análisis visibiliza de qué manera la sociedad de consumo actual, mediante la implantación de determinados espacios físicos, promueve el ideal de felicidad, a la vez que desarrolla nuevas formas de alienación social. El desarrollo de formas de comunicación social, basadas en actitudes y comportamientos caracterizados por la apatía intersubjetiva que se genera en estos espacios, contribuye a consolidar comportamientos mecanizados y (...) deshumanizados. Para verificar todo ello se realiza una investigación etnográfica en El Corte Inglés de Málaga. (shrink)
Este artículo aborda la contextualización de la ética de Butler en un marco teórico que da por supuesta la perspectiva deconstructiva y que, por su parte, es reconstruido de acuerdo a dos criterios, las nociones de “matriz disciplinal” y de “paradigma”. Mediante ellas se trata de proporcionar un sistema de conceptos o categorías usuales en la teoría ética en general, pero redefinidas en términos butlerianos. Estas categorías a las que nos referimos son la acción, los sujetos, los juicios y los (...) criterios morales y, que, aplicadas a la ética de Butler, dan lugar a una ética precaria en la que la humanidad normativa se revela como imposible universalmente y, por tanto, no puede ser utilizada como “fundamento” de un juicio moral universal. Por ejemplo, la que se sugiere mediante las manifestaciones de duelo por la humanidad de “los nuestros”, víctimas de la violencia, de las que queda excluida estructuralmente la humanidad de “los otros”. (shrink)
Este artículo presenta los resultados de la fase de implementación de un estudio de caso que pretende instaurar un modelo de competencias para el desarrollo del pensamiento creativo e innovador a partir de la ingeniería concurrente, como gestión de las prácticas de enseñanza y aprendizaje en una Institución de Educación Superior en Bogotá-Colombia. La actividad se desarrolló bajo un método de investigación acción participativa, el cual buscó explicar razones de los diferentes aspectos que conllevan la planificación, organización y el estudio (...) de un producto o servicio, al intervenir desde un modelo instaurado como proyecto de aula contrastando estos análisis con los primeros resultados de una investigación tipo exploratoria de enfoque cualitativa tomando organizaciones y/o empresas en contextos reales para su diagnóstico de necesidad de innovación. Estos criterios de transferibilidad se basan por una parte sobre una interpretación del enfoque concurrente al realizar el estado del arte existente y los patrones de comportamiento de las empresas para así destacar las dimensiones críticas particulares, por otra parte, se basa en un análisis del tipo de modelo de intervención que permita su implementación en este tipo de empresas. En consecuencia, el modelo de trabajo simultáneo integra las experiencias y la información a través de un conjunto de elementos, que permiten visualizar destrezas creativas para formular ideas novedosas en el ciclo de vida de un producto o servicio, exponiendo la relevancia de la presente estrategia didáctica en integrar la academia con la realidad. (shrink)
En este artículo nos proponemos describir algunos fenómenos lingüísticos que contribuyen a crear la incertidumbre en el texto literario. En particular, analizaremos distintas estrategias narrativas que configuran lo siniestro en un texto que, si bien es un relato de autor, utiliza los modos de decir propios de la leyenda tradicional de transmisión oral: “El Monte de las Ánimas”, de Gustavo Adolfo Bécquer. El efecto inquietante que produce esta leyenda en el lector es resultado de múltiples fenómenos analizados por la teoría (...) literaria. No obstante, queremos proponer un abordaje basado en algunas nociones provenientes del campo de la lingüística y del análisis del discurso como factores determinantes de este efecto. Trataremos tres aspectos: el discurso reproducido o la representación del discurso ajeno, la modalidad epistémica que informa sobre el conocimiento del enunciador frente a los hechos mencionados y los verbos evidenciales que dan cuenta del modo de obtención de la información por parte del locutor, en especial los casos de evidencia indirecta transmitida, el folklore o saber popular. A nuestro entender, la articulación de estos tres elementos constituyen un posible camino de descripción del efecto mencionado. (shrink)
Este artículo es un homenaje a las principales actrices de la escena española, tanto del teatro como del cine. En particular, se fija en la faceta cómica de una serie de artistas que, a través de su talento y su amor a la interpretación, dignificaron su profesión y nos legaron obras memorables. En un mundo predominantemente masculino, estas mujeres lucharon contra las adversidades de su tiempo y supieron labrarse una trayectoria admirable.
Resumen El presente estudio investiga la opción cristiana por la virginidad en el pensamiento de Clemente de Alejandría. En nuestro autor no se encuentra una doctrina organizada sobre la virginidad cristiana, sin embargo, resulta sorprendente que si se estudian estos comentarios aislados de forma conjunta, se puede descubrir de forma clara y certera el pensamiento del maestro de Alejandría sobre la virginidad dentro del cristianismo. En este estudio, las citas textuales de las obras de Clemente de Alejandría servirán de referencia (...) y nos guiarán en el desarrollo de su visión sobre la virginidad cristiana.The present study investigates the Christian option for virginity in the thought of Clement of Alexandria. In our author there is no organized doctrine about Christian virginity, however, it is surprising that if these isolated comments are studied together, we can discover clearly and accurately the thought of the master of Alexandria about virginity within Christianity. In this study, the textual citations of the works of Clement of Alexandria will serve as reference and will guide us in the development of his vision of Christian virginity. (shrink)