What is a larva, if it is not what survives of an ancestor's adult, compressed into a transient pre‐reproductive phase, as suggested by Haeckel's largely disreputed model of evolution by recapitulation? A recently published article hypothesizes that larva and adult of holometabolous insects are developmental expressions of two different genomes coexisting in the same animal as a result of an ancient hybridization event between an onychophoran and a primitive insect with eventless post‐embryonic development. More likely, however, larvae originated from (...) late embryonic or early post‐embryonic stages of ancestors with direct development. Evolutionary novelties would thus be intercalary rather than terminal, with respect to the ancestor's ontogenetic schedule. This scenario, supported by current research on holometabolous insects and marine invertebrates with complex life cycles, offers a serious alternative to the traditional scenario (‘what is early in ontogeny is also early in phylogeny’) underlying the current perception of the evolution of genetic regulatory networks. (shrink)

For over a century, Haeckel's Gastraea theory remained a dominant theory to explain the origin of multicellular animals. According to this theory, the animal ancestor was a blastula‐like colony of uniform cells that gradually evolved cell differentiation. Today, however, genes that typically control metazoan development, cell differentiation, cell‐to‐cell adhesion, and cell‐to‐matrix adhesion are found in various unicellular relatives of the Metazoa, which suggests the origin of the genetic programs of cell differentiation and adhesion in the root of the (...) Opisthokonta. Multicellular stages occurring in the complex life cycles of opisthokont protists (mesomycetozoeans and choanoflagellates) never resemble a blastula. Here, we discuss a more realistic scenario of transition to multicellularity through integration of pre‐existing transient cell types into the body of an early metazoon, which possessed a complex life cycle with a differentiated sedentary filter‐feeding trophic stage and a non‐feeding blastula‐like larva, the synzoospore. Choanoflagellates are considered as forms with secondarily simplified life cycles. (shrink)

Comprehending the origin of marine invertebrate larvae remains a key domain of research for evolutionary biologists, including the repeated origin of direct developmental modes in echinoids. In order to address the latter question, we surveyed existing evidence on relationships of homology between the ectoderm territories of two closely related sea urchin species in the genus Heliocidaris that differ in their developmental mode. Additionally, we explored a recently articulated idea about homology called ‘organizational homology’ (Muller 2003. In: Muller (...) GB, Newman SA, editors. Origination of organismal form: beyond the gene in developmental and evolutionary biology. Cambridge, MA: A Bradford Book, The MIT Press. p 51–69. ) in the context of this specific empirical case study. Applying the perspective of organizational homology to our experimental system of congeneric echinoids has led us to a new hypothesis concerning the ectoderm evolution in these species. The extravestibular ectoderm of the direct developer Heliocidaris erythrogramma is a novel developmental territory that arose as a fusion of the oral and aboral ectoderm territories found in indirect developing echinoids such as Heliocidaris tuberculata. This hypothesis instantiates a theoretical principle concerning the origin of developmental modules, ‘integration’, which has been neglected because the opposite theoretical principle, ‘parcellation’, is more readily observable in events such as gene duplication and divergence (Wagner 1996. Am Zool 36:36–43). J. Exp. Zool. (Mol. Dev. Evol.) 306B:18– 34, 2006. (shrink)

The origin of marine invertebrate larvae has been an area of controversy in developmental evolution for over a century. Here, we address the question of whether a pelagic “larval” or benthic “adult” morphology originated first in metazoan lineages by testing the hypothesis that particular gene co-option patterns will be associated with the origin of feeding, indirect developing larval forms. Empirical evidence bearing on this hypothesis is derivable from gene expression studies of the sea urchin larval gut of (...) two closely related but differently developing congenerics, Heliocidaris tuberculata (feeding indirect-developing larva) and H. erythrogramma (nonfeeding direct developer), given two subsidiary hypotheses. (1) If larval gut gene expression in H. tuberculata was co-opted from an ancestral adult expression pattern, then the gut expression pattern will remain in adult H. erythrogramma despite its direct development. (2) Genes expressed in the larval gut of H. tuberculata will not have a coordinated expression pattern in H. erythrogramma larvae due to loss of a functional gut. Five structural genes expressed in the invaginating archenteron of H. tuberculata during gastrulation exhibit substantially different expression patterns in H. erythrogramma with only one remaining endoderm specific. Expression of these genes in the adult of H. erythrogramma and larval gut of H. tuberculata, but not in H. erythrogramma larval endoderm, supports the hypothesis that they first played roles in the formation of adult structures and were subsequently recruited into larval ontogeny during the origin and evolution of feeding planktotrophic deuterostome larvae. (shrink)

In September of 1869, while studying sponges off the Norwegian island of Gisoe, Ernst Haeckel (1834-1919) discovered a tiny, flagellated ball-shaped organism swimming about in his samples. Appearing first to be the planula larva of an invertebrate marine animal further observation revealed it to be a colony of flagellated cells with a complex life cycle transitioning between multicellular and single-cell stages and several distinct forms of protozoa. Haeckel named it Magosphaera planula (the "magician's ball") and it eventually assumed a central (...) role in his theories of animal evolution, appearing as the modern exemplar of the blastaea stage in his gastraea theory of metazoan evolution. Throughout the latter half of the nineteenth century and into the twentieth it was an object of considerable scientific interest, and yet it was only ever observed by Haeckel himself and then only the once. Eventually it faded altogether from scientific discussion. This paper traces the rise and fall of Magosphaera as an important epistemic object in the theories of Haeckel and other biologists, and an attempt is made to identify what exactly the organism (or organisms!) was that Haeckel observed in the fall of 1869. (shrink)

This volume describes features of biological autonomy and integrates them into the recent discussion of factors in evolution. In recent years ideas about major transitions in evolution are undergoing a revolutionary change. They include questions about the origin of evolutionary innovation, their genetic and epigenetic background, the role of the phenotype, and of changes in ontogenetic pathways. In the present book, it is argued that it is likewise necessary to question the properties of these innovations and what was qualitatively (...) generated during the macroevolutionary transitions. The author states that a recurring central aspect of macroevolutionary innovations is an increase in individual organismal autonomy whereby it is emancipated from the environment with changes in its capacity for flexibility, self-regulation and self-control of behavior. The first chapters define the concept of autonomy and examine its history and its epistemological context. Later chapters demonstrate how changes in autonomy took place during the major evolutionary transitions and investigate the generation of organs and physiological systems. They synthesize material from various disciplines including zoology, comparative physiology, morphology, molecular biology, neurobiology and ethology. It is argued that the concept is also relevant for understanding the relation of the biological evolution of man to his cultural abilities. Finally the relation of autonomy to adaptation, niche construction, phenotypic plasticity and other factors and patterns in evolution is discussed. The text has a clear perspective from the context of systems biology, arguing that the generation of biological autonomy must be interpreted within an integrative systems approach. (shrink)

_This sweeping discussion of the philosophy of evolutionary biology is based on the revolutionary idea that species are not kinds of organisms but wholes composed of organisms._.

First published in 1969. Routledge is an imprint of Taylor & Francis, an informa company.

In his Discourses, Rousseau argues that inequalities of rank, wealth, and power are the inevitable result of the civilizing process. If inequality is intolerable - and Rousseau shows with unparalledled eloquence how it robs us not only of our material but also of our psychological independence - then how can we recover the peaceful self-sufficiency of life in the state of nature? We cannot return to a simpler time, but measuring the costs of progress may help us to imagine alternatives (...) to the corruption and oppressive conformity of modern society. Rousseau's sweeping account of humanity's social and political development epitomizes the innovative boldness of the Englightment, and it is one of the most provocative and influential works of the eighteenth century. This new translation includes all Rousseau's own notes, and Patrick Coleman's introduction builds on recent key scholarship, considering particularly the relationship between political and aesthetic thought. ABOUT THE SERIES: For over 100 years Oxford World's Classics has made available the widest range of literature from around the globe. Each affordable volume reflects Oxford's commitment to scholarship, providing the most accurate text plus a wealth of other valuable features, including expert introductions by leading authorities, helpful notes to clarify the text, up-to-date bibliographies for further study, and much more. (shrink)

By the 1840s and 1850s biogeographical theory had polarized into two opposing views — both of which had their origins in the sixteenth or seventeenth centuries. At issue in this polarization was the question of God's involvement with His creation. At one end of the spectrum were Sclater, Agassiz, Kirby, and others who saw a neatly designed world in which geographical distributions were planned and executed by the hand of God at creation. For most of these naturalists, organisms were created (...) en masse within the regions they now occupy. Disjunct distributions were proof to them that God had indeed created species in situ as many individuals. These naturalists hoped to reveal God's biogeographical plan by discovering His regions of creation. They had hoped to demonstrate a neatly devised set of regions of creation which might be applicable to all creatures, but in attempting to do so, they arrived at conflicting sets of delineations — thus helping to undermine their conceptions of nature in which design (both idealist and utilitarian) played an important part.93At the other end of the biogeographical spectrum were the theoretical ideas of Prichard and Lyell, who viewed a more remote God — one who allowed His creation to be shaped and modified by secondary laws. Lyell in particular wished to leave considerations of design aside, hoping to demonstrate that the shape of the present creation is due to natural laws. Prichard and Lyell saw God's role in the creation of species (and distributions) as being extremely limited. In fact, the regions of creation seen today are in actuality only natural artifacts produced by migrations and barriers. They saw distributions being in constant flux, as was the rest of nature.Those supporting the views of Prichard and Lyell spent a great deal of effort in attempting to remove a major obstacle in their paths — disjunct distributions. If disjunct distributions were indeed the products of separate creative acts, as Sclater and others claimed, then the arguments of Prichard and Lyell would be negated. For if the creation of a species was shown to be the product of multiple creations, then what was the need of migrations and dispersal mechanisms? Also at stake, of course, was the concept of species based upon generation. Darwin was well aware that if the supernatural implications of disjunct distributions could not be refuted, then his evolutionary system — founded upon a species concept based on descent — would be in peril.94A further barrier to the acceptance of the Prichard/Lyell view was the fact that those sympathetic to a nonsupernatural explanation of disjunct distributions could not agree upon a natural explanation for those anomalies, and an internal debate between naturalists within this group raged for decades.95By 1859 a biogeographical stalemate had occurred. Sclater and others, supporting their static view of nature, continued to look for regions of creation, pointing to disjunct distributions in support of their arguments, while those favorable to the views of Prichard and Lyell continued to search for natural explanations for such biogeographical anomalies.The key needed to resolve the biogeographical debate was a credible theory for species origins. By 1858 there were essentially three options for British naturalists: supernatural creation, Lamarckian transmutation, or natural creation. A few British naturalists grasped at these straws, but most workers preferred the option of remaining silent until a more viable explanation for the origin and distribution of species could be advanced.96 And not until the publication of Darwin's theory did that explanation become available. (shrink)

... Difficulty of distinguishing between Varieties and Species — Origin of Domestic ... and Origin— Principle of Selection anciently followed, its Effects— ...

Historians tend to speak of the problem of the origin of species or the species question, as if it were a monolithic problem. In reality, the phrase refers to a, historically, surprisingly fluid and pluriform scientific issue. It has, in the course of the past five centuries, been used in no less than ten different ways or contexts. A clear taxonomy of these separate problems is useful or relevant in two ways. It certainly helps to disentangle confusions that have (...) inevitably emerged in the literature in its absence. It may, secondly, also help us to gain a more thorough understanding, or sharper view, of the history of evolutionary thought. A consequent problem-centric look at that history through the lens of various origin of species problems certainly yields intriguing results, including and particularly for our understanding of the genesis of the Wallace–Darwin theory of evolution through natural selection. (shrink)

Philosophers have not taken the evolution of human beings seriously enough. If they did, argues Peter Munz, many long standing philosophical problems would be resolved. One of philosophical concequences of biology is that all the knowledge produced in evolution is a priori, i.e., established hypothetically by chance mutation and selective retention, not by observation and intelligent induction. For organisms as embodied theories, selection is natural and for theories as disembodied organisms, it is artificial. Following Popper, the growth of knowledge is (...) seen to be continuous from the amoeba to Einstein'. Philosophical Darwinism throws a whole new light on many contemporary debates. It has damaging implications for cognitive science and artificial intelligence, and questions attempts from within biology to reduce mental events to neural processes. More importantly, it provides a rational postmodern alternative to what the author argues are the unreasonable postmodern fashions of Kuhn, Lyotard and Rorty. (shrink)

The essay shows the common ground between music and philosophy from the origin of Western philosophy to the crisis of metaphysical thinking, in particular with Nietzsche and Benjamin. At the beginning, the relationship between philosophy and music is marked by the hegemony of the word on the sound. This is the nature of the Platonic idea of music. With Nietzsche and Benjamin this hegemony is denied and a new vision of the relationship becomes possible. The sound is the (...) class='Hi'>origin both of language and of music. In thinking about this origin, philosophy shows that “thinking about music” is “thinking in music”, and that this thinking is the origin of philosophy itself. (shrink)

Understanding the evolutionary role of environmentally induced phenotypic variation (i.e., plasticity) is an important issue in developmental evolution. A major physiological response to environmental change is cellular stress, which is counteracted by generic stress reactions detoxifying the cell. A model, stress‐induced evolutionary innovation (SIEI), whereby ancestral stress reactions and their corresponding pathways can be transformed into novel structural components of body plans, such as new cell types, is described. Previous findings suggest that the cell differentiation cascade of a cell type (...) critical to pregnancy in humans, the decidual stromal cell, evolved from a cellular stress reaction. It is hypothesized that the stress reaction in these cells was elicited ancestrally via inflammation caused by embryo attachment. The present study proposes that SIEI is a distinct form of plasticity‐based evolutionary change leading to the origin of novel structures rather than adaptive transformation of pre‐existing characters. (shrink)

The institution of the proskynesis at the Roman imperial court seems to have its roots in the world of Iranian peoples, be it the Parthian Empire or Sassanid Persia. Many hints in our sources point eastwards when we consider the origin of this custom. The contacts between the Graeco-Roman and Persian world had a long tradition by the time of Diocletian. For several reasons, it is hard to imagine that the proskynesis, as an act of obeisance before the late (...) antique Emperors, evolved purely within the Empire and unaffected by Eastern influences. (shrink)

In this paper, I show that the idea of logical determinism can be traced back from the Old Babylonian period at least. According to this idea, there are some signs which can explain the appearance of all events. These omens demonstrate the will of gods and their power realized through natural forces. As a result, each event either necessarily appears or necessarily disappears. This idea can be examined as the first version of eternalism – the philosophical belief that each temporal (...) event is actual. In divination lists in Akkadian presented as codes we can reconstruct Boolean matrices showing that the Babylonians used some logical-algebraic structures in their reasoning. The idea of logical contingency was introduced within a new mood of thinking presented by the Greek prose – historical as well as philosophical narrations. In the Jewish genre ’aggādōt, the logical determinism is supposed to be in opposition to the Greek prose. (shrink)

Looks at the life of Charles Darwin, covers the background of the book "On the Origin of Species," presents Darwin's theories and concepts of evolution, and discusses the impact of the book.

After deconstructing the thermodynamic concepts of work and waste, I take up Howard Odum’s idea of energy quality, which tallies the overall amount of energy needed to be dissipated in order to accomplish some work of interest. This was developed from economic considerations that give obvious meaning to the work accomplished. But the energy quality idea can be used to import meaning more generally into Nature. It could be viewed as projecting meaning back from any marked work into preceding energy (...) gradient dissipations that immediately paved the way for it. But any work done by an abiotic dissipative structure, since it would be without positive economic significance, would also be difficult to mark as a starting point for the energy quality calculation. Furthermore, any destructive work as by hurricanes or floods, with negative economic significance, would not seem to merit the quality calculation either. But there has been abiotic work of keen interest to us—that which mediated the origin of life. Some kind of abiotic dissipative structures had to have been the framework that fostered this process, regardless of how it might come to be understood in detail. Since all dissipative structures have the same thermodynamic and informational organization in common, any of them might provide the material context for the origin of something. So we can pick any starting point we wish, and calculate backward what sequence of energy usages would have been necessary to set it up. Given such an open ended project, we could not find an obvious place in any sequence to stop and start the forward the calculation, and so we would need to take it right back to an ultimate beginning, like the insolation of some area, or the outpouring of Earth’s thermal energy. Any energy dissipation might be the beginning of something of importance, and so Nature is as replete with potential meanings as it is with energy gradients. (shrink)

Derrida's introduction to his French translation of Husserl's essay "The Origin of Geometry," arguing that although Husserl privileges speech over writing in an account of meaning and the development of scientific knowledge, this privilege is in fact unstable.

It is at this juncture that post-modernism in the science field lacks that remaining piece of discipline, accepting the existence of an absolute, inner fabric of creative intelligence that is responsible for the vibratory formation of all physical world phenomena. It is for this reason that science alone will not find a viable answer of how the universe was created. Thus far, the science hierarchy has supplied us with theories that are totally incompatible with the belief that the universe had (...) an origin connected to a creative force. Their contradictions come as a result of their atheistic/agnostic beliefs. It is the time in our evolution to make a changing of the guard. We need a new paradigm, a practical application that people can make sense of and find valuable in their daily lives. (shrink)

Are they needed? To be sure. The Darwinian industry, industrious though it is, has failed to provide texts of more than a handful of Darwin's books. If you want to know what Darwin said about barnacles (still an essential reference to cirripedists, apart from any historical importance) you are forced to search shelves, or wait while someone does it for you; some have been in print for a century; various reprints have appeared and since vanished." -Eric Korn,Times Literary Supplement Charles (...) Robert Darwin (1880-1882) has been widely recognized since his own time as one of the most influential writers in the history of Western thought. His books were widely read by specialists and the general public, and his influence had been extended by almost continuous public debate over the last 130 years. New York University Press' edition makes it possible for the first time to review Darwin's public literary output as a whole, plus his scientific journal articles, his private notebooks, and his correspondence. This is the first complete edition containing all of Darwin's published books, featuring definitive texts recording original paginations with Darwin's indexes retained. All illustrations and plates are presented, inclucing 82 color plates of birds and mammals and several folding maps and plates. The set also features a general introduction and index, and textural introductions in each volume. (shrink)

This dissertation is a detailed exploration of Augustine's discussion of the origin of the human soul in the De Genesi ad Litteram. The two central problems addressed are: Why does Augustine abruptly and without explanation abandon his two-phase view of creation and reduce his three hypotheses of the soul's origin to two?, and Why, in spite of what seems to be a preponderance of evidence in favor of the traducianist hypothesis, does Augustine resist it? It is argued that (...) the solution to both of these puzzles is to be found in Augustine's unstated yet persistent commitment to an unmediated account of the soul's origin. It is further argued that if one is sufficiently attentive to the details of Augustine's discussion, it can be seen that Augustine in fact mounts a concerted effort to argue in favor of the creationist hypothesis while simultaneously attempting to block the traducianist hypothesis. (shrink)

Anti-vivisectionists charge that animal experimenters are speciesists people who unjustly discriminate against members of other species. Until recently most defenders of experimentation denied the charge. After the publication of `The Case for the Use of Animals in Biomedical Research' in the New England Journal of Medicine , experimenters had a more aggressive reply: `I am a speciesist. Speciesism is not merely plausible, it is essential for right conduct...'1. Most researchers now embrace Cohen's response as part of their defense of animal (...) experimentation. Cohen asserts that both rights and utilitarian arguments against the use of animals in research fail because they `refuse to recognize the moral differences among species'.2 If we appreciate the profound differences between humans and non-human animals, he says, we would understand why animals do not and could not have rights and why animal pain does not have as much moral weight as human pain. Animal liberationists think speciesism is immoral because they mistakenly equate it with racism and sexism. Cohen claims this equation is `unsound', `atrocious', `utterly specious', and `morally offensive'. Doubtless Cohen is right that the charge of speciesism is founded on an analogy with racism and sexism. He is mistaken, however, in asserting that the comparison is categorically illicit. (shrink)

Philosophers have not taken the evolution of human beings seriously enough. If they did, argues Peter Munz, many long-standing philosophical problems would be resolved. One of the philosophical consequences of biology is that all the knowledge produced in evolution is a priori established hypothetically by chance mutation and selective retention rather than by observation and intelligent induction. For organisms as embodied theories, selection is natural. For theories as disembodied organisms, it is artificial. Following Karl Popper, the growth of knowledge is (...) seen to be continuous from "the amoeba to Einstein." Philosophical Darwinism brings perspective to contemporary debates. It has far-reaching implications for cognitive science and artificial intelligence, and questions attempts from the field of biology to reduce mental events to neural processes. Most importantly, it provides a rational postmodern alternative to what the author views as the unreasonable postmodern theories of Kuhn, Lyotard, and Rorty. (shrink)

Living organisms appear to have agency, the ability to act freely, and humans appear to have free will, the ability to rationally decide what to do. However, it is not clear how such properties can be produced by naturalistic processes, and there are indeed neuroscientific measurements that cast doubt on the existence of free will. Here I present a naturalistic theory of agency, consciousness, and free will. Elementary forms of agency evolved very early in the evolution of life, utilizing an (...) extension of the basic Darwinian scheme that combines and entangles deterministic and stochastic causation. The extension effectively produces an active form of causation, as well as meaning intrinsic to the organism. Consciousness arose when animals evolved advanced nervous systems and social lifestyles that enabled communication mutually affecting each animal’s intrinsic meaning. I argue that various forms of agency that subsequently arose in evolution, preconscious, conscious, intelligent, symbolic, and rational agency, still coexist in humans. This coexistence, combined with the fact that agency is not instantaneous but takes time to build up, makes simple interpretations of neuroscientific results on free will problematic. The conclusion of the study is that conscious free will, including a form of agent causation, is fully consistent with a naturalistic world-view and can be produced by the specific processes discussed. The result closely resembles existing approaches to life and mind that are inspired by phenomenology. (shrink)

In this paper, I show that in the Pāli Canon there was a tradition of Buddhist logic, but this tradition was weak, and the proto-logic we can reconstruct on the basis of the early Pāli texts can be evaluated as a predecessor of the Hindu logic. According to the textual analysis of the Pāli texts, we can claim that at the time of the closing of the Pāli Canon there did not exist the Nyāya philosophy known by the Nyāya Sūtra. (...) Meanwhile, we can assume that the Milindapañha, the best logical source of early Pāli literature, was written under influences of the Gandhāran Buddhists and this text preceded the Nyāya philosophy. (shrink)

Philosophers have not taken the evolution of human beings seriously enough. If they did, argues Peter Munz, many long standing philosophical problems would be resolved. One of philosophical concequences of biology is that all the knowledge produced in evolution is a priori, i.e., established hypothetically by chance mutation and selective retention, not by observation and intelligent induction. For organisms as embodied theories, selection is natural and for theories as disembodied organisms, it is artificial. Following Popper, the growth of knowledge is (...) seen to be continuous from the amoeba to Einstein'. Philosophical Darwinism throws a whole new light on many contemporary debates. It has damaging implications for cognitive science and artificial intelligence, and questions attempts from within biology to reduce mental events to neural processes. More importantly, it provides a rational postmodern alternative to what the author argues are the unreasonable postmodern fashions of Kuhn, Lyotard and Rorty. (shrink)

This book reverses the fundamental tenet of phenomenology-that all consciousness is intentional . Mitscherling rehabilitates the pre-modern concepts of 'intentional being' and 'formal causality' in the construction of a comprehensive phenomenological analysis of embodiment, aesthetic experience, interpretation of texts, moral behavior, and cognition.

This paper accomplishes two tasks. First, I unpack Husserl’s analysis of interest from his 1893 manuscript, “Notes Towards a Theory of Attention and Interest” to demonstrate that it comprises his first rigorous genetic analysis of attention. Specifically, I explore Husserl’s observations about how attentive interest is passively guided by affections, moods, habits, and cognitive tensions. In doing so, I reveal that the early Husserl described attention as always pulled forward to new discoveries via the rhythmic recurrence of tension and pleasure. (...) Second, I demonstrate that “Notes” is the germ of Husserl’s mature genetic phenomenology of attention. The 1893 analysis provides Husserl with all of the philosophical reasons and tools for the construction of his genetic account of attention in his late works. I then discuss how the disclosure of this novel subterranean link can prompt a rethinking of the development of phenomenology. (shrink)

Unified and causal complex-dynamic origin of standard (special and general) relativistic and quantum effects revealed previously at the lowest levels of world interaction dynamics is explicitly generalised to all higher levels of unreduced interaction processes, thus additionally confirming the causally complete character of complex-dynamical, naturally quantised relativity, which does not contain any artificially added, abstract postulates. We demonstrate some elementary applications of this generalised quantum relativity at higher levels of complex brain and social interaction dynamics.